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GRTE-N-1
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Research Report GRTE-N-1
The Elk of Grand Teton and Southern Yellowstone National Parks
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THE STUDY AREA
Vegetation
Scientific and common names for plants mentioned in this bulletin
listed in Appendix I. These
generally follow Davis (1952) or Booth and Wright (1959). Vegetation was
classified during field inspections and by analyzing measurements of
taxa in representative stands. Measurements were by the author
(Appendix II, III, IV) and other workers on the area (Martinka, 1965;
Oswald, 1966; Houston, 1967), following Daubenmire (1959), Parker
(1951), and Cottam and Curtis (1949).
Figure 1 shows the major vegetation types on the area. Table 2 lists
information on their general distribution and the characteristic well as
important associated plants for the more representative stands each
type. Various mixtures of characteristic and important associated plants
occurred in transitional or intermediate successional stages.
Characteristic plants occurred consistently and usually made up the
greater portion of the plant crown cover in representative or
successionally related stands. Important associated plants for other
than forest stands had a crown cover that was slightly less, equal to,
greater than characteristic plants. Important associated plants under
forest canopies made up one-fourth or more of the ground level crown
cover in distinct stands, or had a consistent high frequency of
occurrence in successionally related stands.
Table 2.Classification of vegetation on study area.
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Vegetation Type |
General Distribution |
Characteristic Plants |
Associated Plants |
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BunchgrassShrub |
Residual and glacial till soils on ridgetops and south, east, and west
slopes in valley areas |
Bluebunch WheatgrassDouglas Rabbitbrush Climax
Big SagebrushBitterbrush Stage
Needle and ThreadSandberg Bluegrass Disclimax |
Ricegrass
Rubber Rabbitbrush
Serviceberry
Threetip Sagebrush
Idaho Fescue |
Sagebrush |
Glacial outwash plain, streamcut terraces and alluvial fans |
Big Sagebrush Stage
Big SagebrushBitterbrush Stage |
Bluebunch Wheatgrass
Idaho fescue
Balsamroot
Lupine
Buckwheat
Low Sagebrush |
Valley Meadow |
Alluvial soils along water courses below 7500 feet. |
WillowSedge Stage
Tufted HairgrassSedge Stage
SedgeBluegrass Stage |
Rush
Shrubby Cinquefoil
Silver Sagebrush
Dandelion |
Floodplain Forest |
Stream alluvium up to about 7000 feet. |
Narrowleaf Cottonwood Stage
Blue Spruce Climax |
Russet Buffaloberry
Red Dogwood
Willow
Thinleaf Alder
Balsam Poplar |
Upland Forest |
Glacial till, north and
west slopes within the valley and on a variety of substrates and
exposures in mountain areas up to 10,000 feet. |
Lodgepole Pine Stage
Aspen Stage
DouglasFir Climax
Whitebark Pine Climax
Englemann SpruceSubalpine Fir Climax |
Geyer's Sedge
Fine grass
Arnica
Large Huckleberry
Dwarf Huckleberry
Pachystima
Menziesia
Spirea |
Burn |
Upland forest stands burned since 1930. |
Geyer's SedgeLodgepole Pine Disclimax
Geyer's SedgeSubalpine Fir Disclimax |
(as in Upland Forest) |
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Exceptions to this classification system were the herbland type,
where the crown cover of various forbs was usually greater than the two
characterizing grasses, and the bunchgrass-shrub type, where big
sagebrush/or other shrubs had a greater crown cover than bluebunch
wheatgrass on rocky or more mesic sites. Plant species that
characterized successionally advanced stands which appeared capable of
persisting under the present climate are given a "climax" suffix. Plants
characterizing seral vegetation that could ultimately be replaced by
climax stands are called a "stage." Plants characterizing stands
modified by recent fires or animal use are called a "disclimax." The
classification system represents a modification of criteria used by
Weaver and Clements (1938) and Daubenmire (1953).
Figures 3 and 4 show what appeared to be the general successional
patterns for vegetation in valley and mountain areas. The four types of
climax vegetation below 8,500 feet appeared to develop from a variety of
seral stands on different sites, exposures, and soils. Successional
patterns for subalpine vegetation above 8,500 feet appeared to be
comparatively simple. The dual role of mountain brome, slender
wheatgrass, and tufted hairgrass in climax stands at high elevations and
seral stands at low elevations was evident. Some of associated forbs for
the herbland type probably had similar roles. Dual successional roles
for tree species have been reported by Daubenmire (1953).
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Fig. 3. Plant succession
relationships at elevations from 6200-7500 ft. within valley
areas. (click on image for an enlargement in a new window) |
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Fig. 4. Plant succession
relationships at elevations from about 7000-10,000 ft. within northern
mountain areas. (click on image for an enlargement in a new window) |
Host Douglas-fir and whitebark pine stands on the study area were
considered edaphic climaxes, but on some more mesic sites they were
probably seral to Engelmann spruce and subalpine fir. Ellison (1954)
concluded that scattered stands of Engelmann spruce and subalpine fir
rocky outcrops in the herbland type were seral to herbland. This was on
the Wasatch Plateau in Utah. In this study such stands occurred on rocky
sites in both the herbland and subalpine meadow types and frequently
contained whitebark pine. They were considered to be edaphic climaxes
rather than a seral stage. All lodgepole pine stands were considered
seral, but it is recognized that some stands could be considered "fire
climaxes" as proposed by Daubenmire (1968). Fire and the biotic
influences of wild ungulates in antler rubbing and foraging contributed
to maintaining the forest park type.
The classifications of vegetation and judgments of successional
patterns made during this study should be considered tentative.
Revisions and refinements from more detailed studies are expected. The
general appearance of the major vegetation types is shown on Figure
5.
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Fig. 5a. Approximate center of
Grand Teton Park showing extensive sagebrush type on the outwash plain,
the valley meadow, and bottomland forest types along the Snake River
flood plain, and the upland forest type on glacial moraine
sites. |
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Fig. 5b. The sagebrush type in a
pothole area. |
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Fig. 5c. Valley meadow and
bottomland forest types on Snake River flood plain. |
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Fig. 5d. Valley meadow type on
National Elk Refuge bottomlands and the bunchgrass-shrub type on
adjoining slopes. |
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Fig. 5e. Bunchgrass-shrub type
on south slopes with bottomland developed as a hayfield. |
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Fig. 5f. Herbland type on upper
portions of Chicken Ridge and in the foreground. |
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Fig. 5g. Interspersed forest
park, burn, and upland forest types with herbland above the upper half
of Red Creek Ridge. |
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Fig. 5h. Extensive subalpine
meadow type with scattered Englemman spruce-subalpine fir stands on
Pitchstone Plateau. |
Continued >>>
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