Kyle Joly and caribou
Kyle Joly with a young caribou during migration.
Kyle Joly
Wildlife Biologist
Arctic I&M Network Caribou Vital Sign Lead
4175 Geist Road
Fairbanks, AK 99709

PhD University of Alaska Fairbanks in Wildlife Biology 2011
MS Pennsylvania State University in Ecology (Conservation Biology) 1996
BS Syracuse University in Biology (Population Ecology) 1992

Skills and Interest Areas
Ungulate and carnivore ecology; interactions between wildlife and habitat, including the impacts of wildfire; GPS and remote sensing technology; the nexus between research and management

Professional Experience
2008 - present Wildlife Biologist for Gates of the Arctic NPP and Yukon-Charley Rivers National Preserve and Arctic Network Caribou Vital Sign Lead
2004 - 2008 Wildlife Biologist BLM, Fairbanks, AK
1998 - 2004 Research Wildlife Biologist USGS, Anchorage, AK
1991 - 1998 Biological Technician Alaska, Washington, Colorado

Caribou, Arctic I&M Network Vital Sign
Wolves, Yukon-Charley Rivers National Preserve
Connecting People to Ecosystems--Gates of the Arctic National Park and Preserve


Oster, K. W., P. S. Barboza, D. D. Gustine, K. Joly, and R. D. Shively. 2018. Mineral constraints on arctic caribou (Rangifer tarandus): A spatial and phenological perspective. Ecosphere 9(3):e02160.

Hilderbrand, G. V., D. D. Gustine, B. Mangipane, K. Joly, W. Leacock, L. Stutzman, J. Erlenbach, M. S. Sorum, M. D. Cameron, J. L. Belant, and T. Cambier. 2018. Body size and lean mass of brown bears across and within four diverse ecosystems. Journal of Zoology (in press).

Hilderbrand, G. V., D. D. Gustine, B. Mangipane, K. Joly, W. Leacock, L. Stutzman, J. Erlenbach, M. S. Sorum, M. D. Cameron, J. L. Belant, and T. Cambier. 2018 Phenotypic plasticity across and within brown bear populations in diverse Alaskan ecosystems: Implications for adaptability and resiliency. Polar Biology (in press)

Bowen, L., A. K. Miles, S. Waters, D. Gustine, K. Joly, and G. hilderbrand. 2017 Using gene transcription to assess ecological and anthropological sressors in brown bears. EcoHealth 11 pp. DOI 10.1007/s10393-017-1287-0

Joly, K., T. Craig, M. D. Cameron, A. E. Gall, and M. S. Sorum. 2017. Lying in wait: limiting factors on a low-density ungulate population and the latent traits that can facilitate escape from them. Acta Oecologica 85: 174-183.

Joly, K., J. Rasic, R. Mason, and M. Lukin. 2017. History, purpose, and status of caribou movements in northwest Alaska. Alaska Park Science 17: (in press).

Fullman, T. J., K. Joly, and A. Ackerman. 2017. Effects of environmental features and sport hunting on caribou migration in northwestern Alaska. Movement Ecology 5 (4): 11 pp. DOI 10.1186/s40462-017-0095-z.

Joly, K. 2017. Caribou: Nomads of the North. Alaska Park Science 16 (1): 55-57.

Joly, K., M. S. Sorum, T. Craig, and E. L. Julianus. 2016. The effects of sex, terrain, wildfire, winter severity, and maternal status on habitat selection by moose in north-central Alaska. Alces 52: 101-115.

Guettabi, M., J. Greenberg, J. Little, and K. Joly. 2016. Evaluating potential economic effects of an industrial road on subsistence in north-central Alaska. Arctic 69 (3): 305-317.

Wilson, R. R., L. S. Parrett, K. Joly, and J. R. Dau. 2016. Effects of roads on individual caribou movements during migration. Biological Conservation 195: 2-8.

Joly, K., T. Craig, M. S. Sorum, J. S. McMillan, and M. A. Spindler. 2015. Variation in fine-scale movements of moose (Alces alces) in the upper Koyukuk river drainage, northcentral Alaska. Alces 51: 97-105.

Joly, K., T. Craig, M. S. Sorum, J. S. McMillan, and M. A. Spindler. 2015. Moose (Alces alces) movement patterns in the upper Koyukuk river drainage, northcentral Alaska. Alces 51: 87-96.

Joly, K., S. K. Wasser, and R. Booth. 2015. Non-invasive assessment of the interrelationships of diet, pregnancy rate, group composition, and physiological and nutritional stress of barren-ground caribou in late winter. PLoS One 10 (6): e0127586. doi:10.1371/journalpone.0127586.

Macander, M. J., C. S. Swingley, K. Joly, and M. Raynolds. 2015. Landsat-based snow persistence map for northwest Alaska. Remote Sensing of Environment 163: 23-31.
Joly, K., S. P. Rabinowitch and J. L. Joly. 2015. Dual management of wildlife in Alaska: making federal practice align with federal mandates. George Wright Forum 32: 18-24.

Wilson, R. R., D. D. Gustine, and K. Joly. 2014. Evaluating potential effects of an industrial road on winter habitat of caribou in north-central Alaska. Arctic 67: 472-482.

Bryant, J. P., K. Joly, F. S. Chapin III, D. L. DeAngelis, and K. Kielland. 2014. Can antibrowsing defense regulate the spread of woody vegetation in arctic tundra? Ecography 37: 204-211.

Horne, J. S., T. Craig, K. Joly, G. W. Stout, M. R. Cebrian, and E. O. Garton. 2014. Population characteristics, space use and habitat selection of two non-migratory caribou herds in central Alaska, 1994 – 2009. Rangifer 34 (1): 1-19.

Wilson, R. R., A. Bartsch, K. Joly, J. H. Reynolds, A. Orlando and W. M. Loya. 2013. Frequency, timing, extent, and size of winter thaw-refreeze events in Alaska 2001-2008 detected by remotely sensed microwave backscatter data. Polar Biology 36: 419-426.

Joly, K. Sea ice crossing by migrating caribou, Rangifer tarandus, in northwest Alaska. 2012. Canadian Field-Naturalist 126 (3): 217-220.

Prichard, A. K., K. Joly and J. Dau. 2012. Quantifying telemetry collar bias when age is unknown: a simulation study with a long-lived ungulate. Journal of Wildlife Management 76 (7): 1441-1449.

Joly, K., P. A. Duffy, and T. S. Rupp. 2012. Simulating the effects of climate change on fire regimes in Arctic biomes: implications for caribou and moose habitat. Ecosphere 3 (5): 1-18. Article 36 (

Joly, K. and D. R. Klein. 2011. Complexity of caribou population dynamics in a changing climate. Alaska Park Science 10 (1): 26-31.

Joly, K. 2011. Modeling influences on winter distribution of caribou in northwestern Alaska through use of satellite telemetry. Rangifer Special Issue 19: 75-85.

Joly, K., D. R. Klein, D. L. Verbyla, T. S. Rupp and F. S. Chapin III. 2011. Linkages between large-scale climate patterns and the dynamics of Alaska caribou populations. Ecography 34 (2): 345-352.

Collins, W. B., B. W. Dale, L. G. Adams, D. McElwain, and K. Joly. 2011. Fire, grazing history, lichen abundance, and winter distribution of caribou in Alaska’s taiga. Journal of Wildlife Management 75 (2): 369-377.

Joly, K., F. S. Chapin III, and D. R. Klein. 2010. Winter habitat selection by caribou in relation to lichen abundance, wildfires, grazing and landscape characteristics in northwest Alaska. Écoscience 17 (3): 321-333.

Joly, K., T. S. Rupp, R. R. Jandt, and F. S. Chapin III. 2009. Fire in the range of the Western Arctic Caribou Herd. Alaska Park Science 8 (2): 68-73.

Joly, K., R. R. Jandt, and D. R. Klein. 2009. Decrease of lichens in arctic ecosystems: role of wildfire, caribou and reindeer, competition, and climate change. Polar Research 28 (3): 433-442.

Dale, B. W., L. A. Adams, W. B. Collins, K. Joly, P. Valkenburg, and R. Tobey. 2008. Stochastic and compensatory effects limit persistence of variation in body mass of young caribou. Journal of Mammalogy 89 (5): 1130-1135.

Jandt, R., K. Joly, C. R. Meyers, and C. Racine. 2008. Slow recovery of lichen on burned caribou winter range in Alaska tundra: potential influences of climate warming and other disturbance factors. Arctic, Antarctic, and Alpine Research 40 (1): 89-95.

Joly, K., M. J. Cole, and R. R. Jandt. 2007. Diets of overwintering caribou, Rangifer tarandus, track decadal changes in arctic tundra vegetation. Canadian Field Naturalist 121 (4): 379-383.

Joly, K., P. Bente, and J. Dau. 2007. Response of overwintering caribou to burned habitat in northwest Alaska. Arctic 60 (4): 401-410.

Joly, K., R. R. Jandt, C. R. Meyers, and M. J. Cole. 2007. Changes in vegetative cover on Western Arctic Herd winter range from 1981-2005: potential effects of grazing and climate change. Rangifer Special Issue 17: 199-207.

Joly, K., C. Nellemann, and I. Vistnes. 2006. A reevaluation of caribou distribution near an oilfield road on Alaska’s North Slope. Wildlife Society Bulletin 34 (3): 866-869.

Rupp, T. S., M. Olson, L. G. Adams, B. W. Dale, K. Joly, J. Henkelman, W. B. Collins, and A. M. Starfield. 2006. Simulating the influences of various fire regimes on caribou winter habitat. Ecological Applications 16 (5): 1730-1743.

Joly, K. 2005. The effects of sampling regime on the analysis of movements of overwintering female caribou in east-central Alaska. Rangifer 25 (2): 67-74.

Joly, K., B. W. Dale, W. B. Collins, and L. G. Adams. 2003. Winter habitat use by female caribou in relation to wildland fires in interior Alaska. Canadian Journal of Zoology 81: 1192-1201.

Joly, K., B. W. Dale, W. B. Collins, and L. G. Adams. 2002. Evaluating the impacts of wildland fires on caribou in interior Alaska. Arctic Research of the United States 16: 63-67.

Joly, K., and W. L. Myers. 2001. Patterns of mammalian species richness and habitat associations in Pennsylvania. Biological Conservation 99 (2): 253-260.

Joly, K. 2000. Orphan caribou, Rangifer tarandus, calves; a re-evaluation of overwinter survival data. Canadian Field-Naturalist 114 (2): 322-323.

Myers, W. L., G. P. Patil, and K. Joly. 1997. Echelon approach to areas of concern in synoptic regional monitoring. Environmental and Ecological Statistics 4 (2): 131-152.