The Midden Winter 2022 Issue

Winter 2022 Volume 22 Issue 2
Series of photos showing pinyon juniper encroachment
Images of Grey Cliffs. In the top photo from 1930, note the extensive sagebrush, with pinon pine and juniper trees moving in. In the middle photo from 2008, note the absence of sagebrush, with pinon and juniper now dominating the landscape. The bottom photo from 2022 shows continued growth of PJ.

Top photo: USFS, Middle photo: George Gruell, Bottom photo: NPS

Shifting Baselines

By Bryan Hamilton, Integrated Resource Program Manager

Wait!! What? This wasn’t always a forest?

It is easy to see national parks as static and unchanging. But the world is always moving, and slower, gradual changes often go unnoticed. Over time these changes may become accepted as the status quo, the way things have always been. This phenomenon is called shifting baseline syndrome (Soga and Gaston, 2018).

Shifting baseline describes a gradual change in our accepted norms and expectations for the environment across generations. Our tolerance for environmental degradation increases and our expectations for the natural world are lowered. For example, bison are absent from 98% of their historic range. Yet the functional extinction of bison is often viewed as a conservation success story. Five billion passenger pigeons once darkened skies in eastern North America. It’s impossible for anyone living today to grasp the spectacle and ecological impact of those now extinct flocks. While these examples may seem cliché, distant and even overly dramatic, they show how shifting baselines affect our perception and acceptance of the state of the natural world. In truth similar changes are occurring all around us.

One example is in the vast sagebrush ocean of the Great Basin.

More than any other species, sagebrush defines the Great Basin, forming one of the largest intact habitats in the country. Many animals, like sage grouse and pygmy rabbits, are only found in sagebrush. Sagebrush is also important for recreation and agriculture, carpeting beautiful open spaces with lush springtime wildflowers, vast vistas, and deeply dark night skies. But sagebrush is also one of the most threatened ecosystems in North America.

Fire in the Great Basin is as natural as wind, sun, and rain. Ecosystems here evolved under frequent and low severity fires (Chambers, 2008). Sometimes called “good” fires, many were ignited by lightning, others intentionally started by Native Americans, who used fire as a tool to manage their environment. But colonization virtually eliminated fire as a natural process from the Great Basin, when aggressive fire exclusion began in the 1920’s. This policy had a slow but dramatic effect on sagebrush plant communities.

Without fire, conifers like pinon pine and juniper outcompete and overtake sagebrush. Singleleaf pinon pine and Utah juniper seeds are carried into sagebrush habitat by birds or small mammals, where they establish under a “nurse plant.” Nurse plants provide a cool, moist microclimate, with fertile soil for young trees. Over time these trees “overtop” their sagebrush host, out competing it for critical resources of light and water. Fires historically reset this process every 25-100 years (Knick et al., 2005). In the era of fire exclusion, pinyon juniper woodlands have increased ten fold (Miller and Tausch, 2001), fragmenting the sagebrush ocean into lakes, ponds, and puddles (Welch, 2005).

image showing too much and too little fire
Image showing too much fire on the left and too little fire on the right.

The Nature Conservancy. Illustrator: Kelly Finan, Adapted from: Threat-based land Management: A Field Guide (The SageShare Partnership)

How can we be so certain a site wasn’t always a pinon juniper forest? Or that the site hasn’t reached its natural, stable, climax community like forests in the eastern US? The evidence of this fire driven process is all around us, starting with the very ground we stand on.

Soil formation is not only a geologic process, but a direct reflection of the plants that formed the soils over thousands of years. Based on soil composition, we know that in some forested areas, trees that are present today were not a significant contributor to the creation of those soils. Instead, soil was formed by sagebrush and grasses, not the acidic needles of a conifer forest.

Dendrochronology provides more strong evidence. Pinon and juniper trees can live extraordinarily long lives. The oldest recorded Utah juniper lived nearly 2,000 years; the oldest single needle pinon pine 900 years (Weisberg and Ko, 2012). If the increase in trees is due to fire exclusion, we would expect relatively young trees in former sagebrush habitat. Indeed, almost all trees are less than 100 years old, aligning well with the beginning of colonization and the fire exclusion era. Ancient pinons and junipers are often found in pockets within or adjacent to sagebrush habitat, but these stands are scattered and limited to rocky slopes with thin soils, areas protected from fires. Old growth trees on these sites show regular fire scars, further documentation of the frequency of low intensity fires of the past.

Historic documents and photographs offer still more evidence. Journal entries, place names, wildlife observations, and livestock stocking records indicate a more open, relatively treeless Great Basin as compared to today. Historic photos further corroborate the increase in pinon and juniper and subsequent decrease in sagebrush and grasslands.

Over time, we have forgotten about “good fire” and the historic, open sagebrush landscapes it maintained. We’ve come to view widespread pinyon juniper forests as a permanent part of our environment. But these young forests have major effects on plants and animals. Wildlife, like sage grouse, pygmy rabbits, and yellow-bellied marmots need open sagebrush habitat to survive. Other species like mule deer and migratory birds have declined due to conifer encroachment. Less water is available in streams and springs and there is less native grass and more barren soil. The land is less productive and less diverse.

Fire seemed like the obvious solution to restore sagebrush. But fire itself now presents a challenge. In pinyon juniper forests, dense flammable fuels create catastrophic fires that are hard to control and can damage soils. Following these fires, cheatgrass, an invasive annual, can take over. Cheatgrass is now found on over 17 million acres in the Great Basin and has a negative effect on biodiversity and ecosystem services. Rather than using fire, chainsaws or masticators are used to remove conifers. Sagebrush and other native species, like bluebunch wheatgrass or penstemon, are often seeded on restoration sites after conifer removal. Our success in restoring sagebrush varies but is often better in places with deeper soils, native plants in the understory, and higher precipitation. In addition to restoring sagebrush, conifer removal treatments are designed to recover quickly after wildfires and to resist invasion by cheatgrass. Maintaining sagebrush may require regular maintenance to remove young conifers before they can establish. Restoration projects are carefully monitored to ensure project goals are met.

The sagebrush ocean is the Great Basin. But this vast, beautiful landscape is threatened. Fire exclusion and conifer encroachment have slowly changed sagebrush habitats, leaving these areas vulnerable to catastrophic wildfires and invasion by cheatgrass. As our baselines shift, we may come to see pinyon juniper forests as a fixture, rather than seeing the sagebrush habitat that the conifers have replaced. But active restoration through conifer removal can restore the sagebrush ocean and the wildlife that depend on it.

Chambers, J. C. 2008. Fire and the Great Basin, p. 33-37. In: Collaborative management and research in the Great Basin - examining the issues and developing a framework for action.
J. C. D. Chambers, Nora; Evenden, Angela (ed.). Gen. Tech. Rep. RMRS-GTR-204. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station.
Knick, S. T., A. L. Holmes, and R. Miller. 2005. The role of fire in structuring sagebrush habitats and bird communities. Studies In Avian Biology. 30:1-13.
Miller, R. F., and R. J. Tausch. 2001. The role of fire in pinyon and juniper woodlands: a descriptive analysis. Proceedings of the Invasive Species Workshop: the Role of Fire in the Control and Spread of Invasive Species. Fire Conference 2000: the First National Congress on Fire Ecology, Prevention, and Management. Miscellaneous Publication No. 11, Tall Timbers Research Station, Tallahassee, FL.:15-30.
Soga, M., and K. J. Gaston. 2018. Shifting baseline syndrome: causes, consequences, and implications. Frontiers in Ecology and the Environment. 16:222-230.
Weisberg, P. J., and D. W. Ko. 2012. Old tree morphology in singleleaf pinyon pine (Pinus monophylla). Forest Ecology and Management. 263:67-73.
Welch, B. L. 2005. Big sagebrush: A sea fragmented into lakes, ponds, and puddles. Gen. Tech Rep. RMRS-GTR-144. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station:210.

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cave drawing
Potential cattle brand symbol made using carbide in 1941.

Brianna Patterson

Hidden Stories of Snake Creek Cave

By Brianna Patterson, Archaeologist, Great Basin Institute

For thousands of years, humans have entered caves and made their mark. From rock writing to historic inscriptions, these cultural resources capture a moment of the composer’s life. Such is the case for the more than 200 historic inscriptions within Snake Creek Cave in Great Basin National Park. Archaeologists see these entries as more than just names and dates. They use the information left behind to piece together past lives. It’s the stories archaeologists tell that are at the heart of our work.

A small crew composed of volunteers and one archaeologist recently started uncovering the history of Snake Creek Cave’s former explorers. Within three hours, the crew had recorded 104 inscriptions, which cover a span of time from 1885 to 1978. These inscriptions include several fascinating entries such as “Kenneth B. Vanda H. 1932 Sweethearts”, “Francis April 21, 1935 3rd Grade”, and a potential cattle brand symbol made using carbide.
cave writing
Percy and Bert Loper inscriptions from July 10, 1931.

Brianna Patterson.

Though there is still much work left, the crew’s efforts have already made a connection to a man named Bert Loper. Through genealogical research, the crew discovered Bert was born in 1913 in Burbank, Utah a little over twenty miles from the site. In 1930, he first entered the cave at seventeen-years-old and returned the next year with his thirteen-year-old brother, Percy Loper. In 1935, Bert moved to Las Vegas, Nevada to work as a steel cleaner and painter at the Hoover Dam. Sadly, Bert died on November 17th after roughly three months on the job. He was electrocuted while working on the rails of a crane in the power house and fell sixty feet.

The Loper family didn’t return to Snake Creek Cave until ten years after Bert’s death. Fifteen-year-old Lavern Loper explored the cave in 1945, and seventeen-year-old Guy Loper ventured there in 1946. Both boys left their inscriptions just as their older brothers had. In 1951, Percy Loper returned to the cave twenty years after his first visit with Bert. It’s possible the cave and Bert’s inscriptions served as a memorial for his siblings, a place where they could remember and feel connected to their brother.

From just a few names and dates, the crew has started to piece together one family’s bond to Snake Creek Cave. The Lopers' inscriptions and other links to the past must be preserved by leaving these cultural resources as they are. This is true for the cave as well. Caves form over millions of years, and both historic and modern inscriptions permanently ruin this important natural resource. While we can’t undo the damage from historic inscriptions, we can prevent further harm to the cave by leaving it as it is.

We hope the research conducted in Snake Creek Cave will continue to reveal hidden stories about the cave’s visitors. This work is associated with a partnership between the Great Basin Institute, Great Basin National Park, and Humboldt-Toiyabe National Forest and is funded by the Southern Nevada Public Lands Management Act. The goal is to study wild caves within White Pine County to better understand and manage them. Thank you to the volunteers who followed a stranger into a dark hole to help record historic inscriptions. And extra thanks to the volunteer who immediately started researching and found the mysterious Bert Loper.

high elevation plants
Focal species in clockwise order from top left: Nachlinger catchfly, Holmgren’s buckwheat, Pennel beardtongue, and Nevada primrose.

Wade Plafcan

Life at the Top: Range Shifts of Four High Elevation Plants

By Wade Plafcan and Thomas Albright, Researchers, University of Nevada, Reno

Starting in fall of 2021, Great Basin National Park, the University of Nevada, Reno, and botanist Jan Nachlinger began a project to track changes in recent and future distributions of four rare alpine plant species.

Nachlinger catchfly (Silene nachlingerae), Holmgren’s buckwheat (Eriogonum holmgrenii), Nevada primrose (Primula cusickiana var. nevadensis), and Pennell beardtongue (Penstemon leiophyllus var. francisci-pennellii) (Figure 1) are all rare, herbaceous plant species of conservation concern. They occur in high elevation habitats and are endemics, meaning they are limited to Great Basin National Park or a few other ranges in eastern Nevada. Being limited to the highest elevations means that there is no room for them to move higher to escape increasing temperatures and having a limited endemic distribution means they don’t have ‘backup’ locations. This combination makes these species especially vulnerable to anthropogenic driven climate change.

To investigate these changes we mapped the recent, current, and predicted future distributions of these plants using two surveys separated by 15 years. Our primary objective was to detect changes in plant distribution and elevation a) between two surveys conducted in 2004-2006 and 2021, and b) between current and projected mid-21st century climates. Additionally, we wanted to determine which environmental factors are most associated with plant habitat and identify locations important for conservation.

three people standing on top of mountain
The veg crew works up in the alpine.

Wade Plafcan

With GPS units, sturdy boots, and lots of water, our team surveyed thousands of acres for plant locations in the summers of 2021 and 2022. On top of this, we were lucky that some forward-looking naturalists had conducted extensive surveys in 2004-2006. This gave us the opportunity to look for changes: places where the plants may be dying off and locations where new populations may be establishing. Overall, surveyors found about half of the plants occupying the same locations from the previous survey. This is an impressive survey effort, as finding a single plant sometimes takes 3-4 people on their hands and knees for several minutes! A comparative analysis between the two surveys revealed a potentially alarming trend. For all species, individuals found lower in the species’ elevational range were more likely to be absent, while the new and persistent populations occurred higher in elevation. This suggests an upward elevational range shift.

Research indicates many alpine species are moving up in elevation to track more suitable (often cooler) habitat conditions. Unfortunately, for species already occurring on the tops of mountains, they may ‘run out of real estate.’ For example, the new populations of Holmgren’s buckwheat occurred at a mean elevation of 11,150 ft compared to 10,500 ft mean elevation of the individuals lost since the previous survey. A vertical difference as tall as the Space Needle!

habitat maps for alpine plants
Example Species Distribution Model map for Nachlinger catchfly: range size change map showing year changes into 2050/2070 with four different climate projections. Colors indicate habitat changes with red indicating loss, yellow being stable, and green areas of gain. Black line is NPS boundary.

But what about conserving these species into the future? The best way to ensure the preservation of species vulnerable to human-driven climate change is to reduce the emissions of greenhouse gasses globally. Beyond those measures, an effective conservation plan for rare species needs to identify locations of suitable habitat to prioritize and target conservation efforts. Species distribution models (SDMs) give insight into these locations and allow for prediction of current and future habitat. SDMs relate a species’ geographic location with environmental variables to produce predictive distribution maps. Plant locations were matched with climate, topography, and soil variables. Through this process we were able to quantify which environmental variables were important for each species. Current habitat maps are useful for finding new populations and as an estimate of total current habitat.

To understand what might happen in the future, SDMs were projected into years 2050 and 2070. Multiple climate projections were used to hedge our bets on the uncertainty of future conditions. These projections included combinations of different amounts of greenhouse gas emissions, warming and both increasing and decreasing precipitation.

Results varied by species and by climate projection. Nachlinger catchfly is of most conservation concern, showing a potential loss of almost all favorable habitat by 2050. Pennell beardtongue may also lose a moderate amount of habitat and has the largest predicted increase in mean elevation, which could lead to habitat fragmentation over time. Holmgren’s buckwheat and Nevada primrose are predicted to be relatively stable and may even expand their habitat under some climate projections. However, contrary to the future models, the survey comparison results show these species may be experiencing upward range shifts already. Only time will reveal which method and analysis is more accurate.

Continued monitoring of these species will be important to track changes and verify model predictions. Monitoring climate change-altered precipitation levels will be of paramount importance, as the driest climate projection predicts substantial habitat loss for all species. Snow, precipitation, and temperature are very important variables for the biology and phenology of these species. Fortunately, the SNOTEL (Snowpack Telemetry) weather station, established in 2011 near Wheeler Peak, will provide great data on changes associated with these climate variables.

Our finding that range shifts from a warming climate may already be occurring is a bit jarring but not unexpected. Our hope for the future is that we can avoid the worst of climate change by reducing emissions globally, and that conservation biologists can help to ensure that species and habitats are protected in the right places to be able to weather the climate changes that do come our way.

person holding cutthroat trout in water
A Bonneville cutthroat trout in one of the Park's small streams

NPS Photo/Joey Danielson

Genetic Analysis of West Desert Bonneville Cutthroat Trout

By Joseph Danielson, Biological Science Technician

Great Basin National Park (GRBA) is currently working in collaboration with Brigham Young University (BYU) to determine genetic variation and purity among Bonneville Cutthroat Trout (BCT) populations in Nevada.

The BCT is a subspecies of cutthroat trout that lived throughout the Bonneville Basin, which included Lake Bonneville (now the Great Salt Lake) and its tributaries, at the end of the last ice age. Once thought to be extinct, genetically pure populations have been found in recent decades in Utah, Idaho, Wyoming, and Nevada. The BCT is the only species of trout native to GRBA and east central Nevada. Careful management has allowed the BCT to remain off the endangered species list, but human encroachment, the introduction of non-native trout, an increase in fire severity, and worsening drought still threatens the species, particularly at the periphery of its range.

The information gained through genetic analysis will be used to determine the source population(s) for the Baker Lake BCT introduction and for other future reintroductions and management actions. Current BCT populations in the region are isolated and rather small in numbers, particularly with the recent drought years, which can negatively affect the genetic diversity of these populations through inbreeding depression and genetic drift. Identifying these genetically diminished populations, and whether certain streams could benefit from inter-population translocations will help us maintain healthy populations of fish.
two biologists with a fish and bucket
Great Basin National Park biologists collect a fin clip.

NPS photo/Joey Danielson

To aid in the preservation of this species, the Great Basin National Park fisheries crew is working to restore habitat and protect BCT in local streams. However, threats to, and recent loss of several populations due to fire has prompted additional proactive measures such as the high elevation lake refugia project in which we are moving BCT into Baker and Johnson Lakes to increase the number of populations in Nevada. For a successful fish move, we need to identify populations which have enough fish and genetic diversity to be used as a source population, so we have been collecting genetic samples from BCT throughout their range in Nevada. We do this by clipping a small portion of the caudal fin of at least 30 fish per stream. So far, we have sampled approximately 500 fish from 15 out of the 16 streams in Nevada currently occupied by the species.

BYU’s analysis will shed light on how much genetic variation there is in fish within and among Nevada’s BCT streams and show if there has been any introgression with non-native trout species. Although historically BCT were the only trout species native to east central Nevada, the haphazard introduction of other trout species following white European settlement has led to BCT being either outcompeted and/or hybridized in many areas. Locally, the Rainbow Trout is of particular concern as it can successfully spawn with BCT to produce fertile hybrid offspring and has done so in GRBA streams in the past.

Nevada’s populations of BCT were reestablished using few local source populations and we expect that the genetic analysis will reflect these different lineages. While these populations at the time were not considered to be bottlenecked, the size of the populations suggest that genetic diversity could be an issue in the future. Once the genetic analysis is complete, the Park will have information needed to proceed with introducing BCT into Baker Lake in 2023 as well as make decisions on how to optimally manage all the isolated BCT populations in Nevada.
Small harvestman with green paint on it
Green paint was used in September to mark harvestmen for a mark and recapture study of these cave dwellers to learn more about them.

Jean Krejca

Mark and Recapture Study for the Model Cave Harvestman, Sclerobunus ungulatus

By Shiloh McCollum, Biologist, Great Basin Institute

Arachnids, particularly shy cavernicolous ones, keep their secrets close to their scute. In fact, they don’t like to spill the beans at all. In order to engage them in disclosing all their secrets we cannot whisper sweet nothings, instead hard science is called to the field of battle.

Harvestmen are solitary, photophobic omnivores that can sometimes exceed, in biomass, the number of spiders1. Cavernicolous species often have less seasonal variation than surface species1, but overall, not a lot is known about them. Sometimes they have continuous reproduction, sometimes not. Sometimes they guard their eggs and juveniles, sometimes not. Sometimes they are highly vagile (like to move), sometimes not.1

Sclerobunus ungulatus, the Model Cave Harvestman, is endemic to Great Basin Caves and even less is known of their secrets. We do not know if they feed on feces or springtails1. We do not know much about their biology, population dynamics, vagility, or really any of their secrets. We do know that the type specimen was collected in 1952 from Model Cave2. It differs from the closely related Sclerobunus cavicolens by a variety of features, including a lack of pigmentation2. It also differs from Sclerobunus madhousensis by a longer second leg and a limited range (S. madhousensis is endemic to caves near Provo, UT)2. Its current range covers the Baker Creek Watershed and 2 high elevation caves3,4. Thus, we are introducing a mark and recapture study in order to gain more knowledge and plunder secrets like mad pirates… or mad scientists (really, a very fine line sometimes).
tape measures in cave
Defining transects in Model Cave

Jean Krejca

Using a similar methodology to a Brazilian Opiliones study1, we determined to make 4 transects in Model Cave of 3 meters in length.

The study is conducted every month by the same two researchers to reduce variables. Each researcher searches a transect for 15 minutes, collects all the specimens of interest they find, and makes observations of other species nearby (possible prey items). Then the other researcher does the same for another 15-minute pass. Once captured, the specimens are marked with fluorescent, acrylic paint with unique identifiers. A different color is used for each capture date, and a different leg is painted based on the transect.
harvestman with pink paint on it
In October, pink paint was used to mark harvestmen.

Shiloh McCollum

Once marked, specimens are photographed for size/age class information using a microscale and to record the unique identification of the specimen. All harvestmen are returned to the transect from which they were collected.

After a year of our study, data will be compiled and crunched using appropriate statistical methods1 to learn as much as we can about population size, size classes, life span, reproduction, and any other biological observations we can glean during our study. Using Model Cave, we can then begin to unravel S. ungulatus’ secrets.


  1. Augusto Macedo Mestre L, Pinto-da-Rocha R(2004) Population Dynamics of an Isolated Population of the Harvestmen Ilhaia cuspidata (Opiliones, Gonyleptidae) in Araucaria Forest (Curitiba, Parana, Brazil). The Journal of Arachnology 32:208-220
  2. Derkarabetian S, Hedin M (2014) Integrative Taxonomy and Secies Delimination in Harvestmen: A revision of the Western North American Genus Sclerobunus (Opiliones: Laniatores: Travunioidea). PLoS ONE 9(8): e104982
  3. Krejca JK, Taylor SJ (2003) A Biological Inventory of Eight Caves in Great Basin National Park. White Paper. Park Files.
  4. Taylor SJ, Krejca JK, Slay ME (2008) Cave biota of Great Basin National Park, White Pine County, Nevada. Illinois National Historical Survey, Champaign, Illinois. Center for Biodiversity Technical Report 25: 398 pgs.
photos of a pretty pink flower
Are these different primrose varieties or not? What is currently known as Primula cusickiana: A: var. maguirei in the Bear River Range of Utah, B: var. cusiciana Snake River Plain and Jarbidge, NV, C: var. domensis in the House Range in Utah, and D: var. nevadensis in the Snake and Grant Ranges.

Austin Koontz

A Closer Look at Nevada Primrose

By Austin Koontz, Researcher

At the summit of Mt. Washington grows a rare plant with a unique backstory. Primula cusickiana var. nevadensis, the Nevada Primrose, is a perennial plant with showy purple petals (Panel D in image to the right), and was first described as its own species in 1967 by Noel Holmgren. It is part of the Primula cusickiana species complex: a group of related plants with similar morphologies found throughout the Great Basin. The other varieties in the complex are:
  • P. cusickiana var. maguirei, in the Bear River Range of Utah (A);
  • P. cusickiana var. cusickiana, across the Snake River Plain in Idaho but with a population also in Jarbidge, Nevada (B)
  • P. cusickiana var. domensis, in the House Range in Utah, just across the border from Great Basin National Park (C)
While a Master’s student at Utah State University from 2018-2020, I conducted a genetic survey of these four varieties under the instruction of Dr. Paul Wolf and Dr. Will Pearse. The goal of our research was to determine how genetically distinct these rare plants are, and if they are properly described as “varieties” of the same species.
map of Nevada, Utah, and Idaho showing where Primula cusickiana occurs
Distribution map of the primrose Primula cusickiana. The different colors distinguish different genetics.
Our findings suggest not! We found very strong divisions between the isolated populations of this plant, suggesting that each is undergoing “cryptic speciation”: a process by which individuals in different locations look very similar, but are genetically very distinct. The history of the Great Basin may provide some clues as to what’s driving this process.

At the end of the Pleistocene 11,700 years ago, the Great Basin was a cool, relatively wet area featuring large lakes, such as the ancient Lake Lahontan and Lake Bonneville. At this time, the ancestor of Primula cusickiana was likely prevalent throughout the region. Over the course of the next 10,000 years (the Holocene), the Great Basin became hotter and drier. Mountaintops, however, continued to provide the cool conditions many organisms were used to; as the climate changed, these organisms retreated to these mountainous regions, which scientists call refugia. Primula was one of these organisms: as it retreated towards higher elevations to follow the cool, moist soils it prefers, its populations became fragmented. This fragmentation led to populations becoming more and more genetically unique, leading to the strong genetic divisions we see between populations today.

Our research also shined a light on the unique nevadensis population on Mt. Washington. Our research suggests that this population of nevadensis is a hybrid between the domensis population (to the east in the House Range) and another nevadensis population further south (in the Grant Range). More needs to be done to characterize these two populations, but our work suggests that the Mt. Washington population (and indeed all populations) is very unique!

I hope this work is built upon and can lead to the protection of these rare and unique plants! Interested readers can find out more about this work in the article here, published in Systematic Botany.
single tree in orchard
Historic Moorpark variety apricot tree in Lehman Orchard

NPS Photo

Preserving Historic Lehman Orchard

by Meg Horner, Biologist

Lehman Orchard changed for the better this past summer. With funding from the Southern Nevada Public Lands Management Act (SNPLMA) Conservation Initiative, park staff replaced the irrigation system, planted new fruit trees, installed new fencing, and designed and installed informational signs and a viewing platform. The upgrades and orchard restoration work completed this year will help preserve Lehman Orchard for years to come.

The history of Lehman Orchard is lengthy and not well documented. The orchard was likely planted in the 1890s soon after Absalom S. Lehman settled near Lehman Caves and started promoting the caves as a tourist attraction. Records indicate Lehman planted fruit trees and constructed an aqueduct that brought water from Lehman Creek and nearby springs to irrigate the orchard. Four kinds of fruit trees were planted in the original orchard including apricot, apple, pear, and peach. Fruit from the orchard became an important source of food for the local community and income for Absalom Lehman and his family. Remarkably, a few of the apricots planted over a century ago are believed to be the same apricots still growing in the orchard today.
Black and white photo of small fruit trees
Part of the historic Lehman Orchard circa 1930

NPS photo

In 1922, Lehman Caves National Monument was designated as a new monument without mention of the orchard or aqueduct. By the 1930s, the condition of many of the fruit trees had deteriorated. The Civilian Conservation Corps removed an unknown number of trees in 1934. By 1940, there were 18 trees remaining in the orchard (Superintendent Report, January 1943). Additional trees were removed for construction projects in the 1940s, 50s, and 60s and made using the aqueduct for irrigation no longer possible. The Lehman Orchard and Aqueduct were added to the National Registry of Historic Places in 1975 because of their importance as a food source and an example of late 19th and early 20th century farming and ranching in eastern Nevada. In the two decades that followed the nomination, the National Park Service (NPS) began work to revitalize the orchard by planting new trees and adding irrigation and fencing. Additional trees were planted outside the historic Lehman Orchard near the Lehman Caves Visitor Center and at the Forest Service Ranger Station in the town of Baker.

The most recent orchard restoration and preservation efforts began in 2017 with funding from the SNPLMA program as part of the Protection, Stabilization, and Restoration of the Lehman Caves Historic Area Project. Park staff researched the history of Lehman Orchard as well as the species and varieties of fruit trees growing in the local area. A greenhouse was constructed to grow seedlings for future planting. Seeds and cuttings were collected from the historic apricots in Lehman Orchard; Marianna root stock was purchased and grafted with ‘volunteer’ apples that were discovered several years earlier growing at two historic camp sites along Baker and Lehman Creeks; and Elberta variety heirloom peaches were purchased. Pear cuttings were collected from a local ranch, but seedlings did not survive.
view of trees and pullout
Lehman Orchard with historic apricots on the left and viewing platform and new seedlings on the right. Taken November 2022.

NPS Photo/Meg Horner

Twenty-two new fruit trees (apricots, peaches, and apples) were planted into Lehman Orchard in 2022, the 100-year anniversary of Lehman Caves National Monument. A new, water-wise irrigation system was installed before planting to ensure consistent water for the seedlings plus the 15 trees already established in the orchard – historic apricots and peaches, apples, and apricots planted in the 1980s and 90s. Leaf samples from historic apricot and apple trees were collected for genetic analysis. Finally, interpretive signs and a viewing platform were installed that describe orchard maintenance and pruning methods and invite visitors to relax and enjoy the view.

Initial results from genetic testing solved a couple mysteries about the heirloom varieties of Lehman Orchard’s historic trees. One of the historic apricot samples was a match for Moorpark, a very old and highly regarded apricot variety dating back to the 1600s when the trees were brought from China to England for cultivation. Several of the other historic apricots in Lehman Orchard were likely seedlings from a Moorpark variety. There were no matches for the apples that were used for grafting from Baker or Lehman Creek. Further testing is planned to identify additional varieties and help preserve the historic and genetic character of Lehman Orchard. The NPS is committed to the preservation of Lehman Orchard, the legacy of the century-old apricots that still produce fruit every few years and sharing the story of this unique resource for another 100 years.

Selected Publications about the Park

  • Hankin, L. E. (2022). Persistence potential of Great Basin high-elevation conifers under novel climates and disturbance regimes (Doctoral dissertation). Link
  • Koontz, A., Pearse, W. D., & Wolf, P. G. (2022). Pronounced genetic separation among varieties of the Primula cusickiana species complex, a Great Basin endemic. Systematic Botany 47(3): 887-897. Link
  • Plafcan, W. (2022). Recent and Future Climate-Induced Range Shifts of Four High-Elevation Rare Forbs in Great Basin National Park, USA (Doctoral dissertation, University of Nevada, Reno). Link
  • Walter, L. M. (2022). Reconstructions of Late Pleistocene Mountain Glacier Equilibrium Line Altitudes and Paleoclimate in the Great Basin (Doctoral dissertation, North Dakota State University). Link

Upcoming Events

  • December 14: Christmas Bird Count: Snake Valley Help gather data during this fun citizen science project! Contact the park ecologist for more info.
  • December 15: Christmas Bird Count: Ely Two bird counts, two days! Contact Nancy Herms at 775-289-1800 for more info.
  • January 20-22: Lehman Caves Lint and Restoration Camp. Pick lint, remove spent carbide, vacuum trails, and more while you show the cave some love! e-mail us for more info.
packrat with headlamp and backpack reading a paper
This woodrat knows of the many treasures to be found in middens!

Illustration by Emily Hale

More information

The Midden is the Resource Management newsletter for Great Basin National Park. A summer and winter issue are posted each year. We welcome submissions of articles or drawings relating to natural and cultural resource management and research in the park. They can be sent to: Editor, Resource Management, Great Basin National Park, Baker, NV 89311 Or call us at: (775) 234-7331.

Superintendent: James Woolsey
Integrated Resource Program Manager: Bryan Hamilton
Editor & Layout: Gretchen Baker

What’s a midden?

A midden is a fancy name for a pile of trash, often left by pack rats. Pack rats leave middens near their nests, which may be continuously occupied for hundreds, or even thousands, of years. Each layer of trash contains twigs, seeds, animal bones and other material, which is cemented together by urine. Over time, the midden becomes a treasure trove of information for plant ecologists, climate change scientists, and others who want to learn about past climatic conditions and vegetation patterns dating back as far as 25,000 years. Great Basin National Park contains many middens.
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Last updated: November 9, 2022

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Contact Info

Mailing Address:

100 Great Basin National Park
Baker, NV 89311


Available 8:00 am - 4:00 pm, Monday through Friday. Closed on Thanksgiving, Christmas, New Year's Day

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