Carol McIntyre - Wildlife Biologist

A woman sits outside on a rock with her dog sitting at her feet.
Carol McIntyre is a wildlife biologist at Denali National Park and Preserve.

Carol McIntyre
Wildlife Biologist
Denali National Park and Preserve
4175 Geist Road
Fairbanks, Alaska 99709 United States

Carol McIntyre is a Wildlife Biologist with Denali National Park and Preserve, Alaska. She started studying the ecology of Golden Eagles and other birds in Denali in 1987. She has published the results of her studies in scientific journals and co-authored the Birds of the World Golden Eagle species account and the book, Birds of Denali. Carol was elected a Fellow of the American Ornithologists' Union in 2016 and presented with the Raptor Research Foundation’s Fran and Frederick Hamerstrom Award in 2021. When she isn’t in the field studying birds, she's out on the trail with her husband Ray and their small team of very large Alaska sled dogs.

Check out this story from the Audubon Society about How Carol McIntyre Became a Rock Star Among Raptor Biologists.

Critical Connections
Golden Eagles

PhD Oregon State University, Wildlife Science
MS University of Alaska-Fairbanks, Wildlife Management
BS East Stroudsburge University

Areas of Interest

  • Population ecology of northern breeding birds.

  • Conservation of areas and resources used by northern breeding birds across their annual cycle.

  • Response of northern breeding birds to cascading effects of climate change.


Mizel, J. D., J. H. Schmidt, C. A. Roland, and C. L. McIntyre. 2023. Tree and shrub expansion at treeline drive contrasting responses in a subarctic passerine community. Journal of Animal Ecology 92(6): 1256-1266.

Adams, E. M., I.J. Stenhouse, A. T. Gilbert, J. Boelsma, G. Gress, C. S. Weidensaul, C. Grigsby, E. J. Williams, L. Phillips, and C. L. McIntyre. 2022. The first documentation of the Nearctic-Paleotropical migratory route of the Arctic Warbler. Ecology and Evolution 12(9): e9223.

McIntyre, C. 2021. An introduction to some of the high-flying technology used to study the movements of Alaska’s migratory birds. Alaska Park Science 20(1): 74-83.

Mizel, J.D., J. H. Schmidt, and C. L. McIntyre. 2021. Climate and weather have differential effects in a high-latitude passerine community. Oecologia.

Davidson, S. C., G. Bohrer, E. Gurarie, S. LaPoint, P. J. Mahoney, ... K. Joly, ... J. P. Lawler, ...B. Mangipane, ... C. L. McIntyre, ... P. A. Owen, ... M. S. Sorum, ... et al. 2020. Ecological insights from three decades of animal movement tracking across a changing Arctic. Science 370(6517): 712-715.

Franke, A., K. Hawkshaw, S. Ambrose, ... C. L. McIntyre, et al. 2020. Status and trends of circumpolar peregrine falcon and gyrfalcon populations. Ambio 49(3): 762-783.

Katzner, T. E., M. N. Kochert, K. Steenhof, C. L. McIntyre, E. H. Craig, and T. A. Miller. 2020. Golden Eagle (Aquila chrysaetos), version 2.0. In Birds of the World (P. G. Rodewald and B. K. Keeney, editors). Cornell Lab of Ornithology, Ithaca, NY, USA.

Stenhouse, I. J., E. M. Adams, L. M. Phillips, S. Weidensaul, and C. L. McIntyre. 2019. A preliminary assessment of mercury in the feathers of migratory songbirds breeding in the North American subarctic. Ecotoxicology, DOI 10.1007/s10646-019-02105-2.

Mizel, J. D., Schmidt, J. H., Phillips, L. M. and Mcintyre, C.L., 2019. A binomial N‐mixture model for estimating arrival and departure timing. Methods in Ecology and Evolution 10(7): 1062-1071.

DeLuca, W. V., B. K. Woodworth, S. A. Mackenzie, A. E. M. Newman, H. A. Cooke, L. M. Phillips, N. E. Freeman, A. O. Sutton, L. Tauzer, C. McIntyre, I. J. Stenhouse, S. Weidensaul, P. D. Taylor, and D. R. Norris. 2019. A boreal songbird’s 20,000 km migration across North America and the Atlantic Ocean. Ecology 00(00):e02651. 10.1002/ecy.2651

Eisaguirre, J. M., T. L. Booms, C. P. Barger, C. L. McIntyre, S. B. Lewis, and G. A. Breed. 2018. Local meteorological conditions reroute a migration. Proceedings Royal Society B 285:20181779.

Mizel, J. D.,C.L. McIntyre, S.B. Lewis, M.S. Lindberg, and J.H. Schmidt. 2018. A multi‐state, time‐removal model for population dynamics of cliff‐nesting raptors. Journal of Wildlife Management 82:1701-1710. doi:10.1002/jwmg.21526

Fuglei, E., J. A. Henden, C. T. Callahan, O. Gilg, J. Hansen, R. A. Ims, A. P. Isaev, J. Lang, C. L. McIntyre, R. A. Merizon, O. Y. Mineev, Y. N. Mineev, D. Mossop, O. K. Nielsen, E. B. Nilsen, A. O. Pedersen, N. M. Schmidt, B. Sittler, M. H. Willebrand, and K. Martin. 2018. Circumpolar status of Arctic ptarmigan: Population dynamics and trends. Ambio DOI 10.1007/s13280-019-01191-0.

McIntyre, C.L., and S.B. Lewis. 2017. Statewide movements of non-territorial Golden Eagles in Alaska during the breeding season: information for developing effective conservation plans. Alaska Park Science 17:

Steenhof, K., M.N. Kochert, C.L. McIntyre and J.L. Brown. 2017. Coming to terms about describing golden eagle reproduction. Journal of Raptor Research 51: 378-390.

Brown, J.L., B. Bedrosian, D.A. Bell, M.A. Braham, J. Cooper, R.H. Crandall, J. DiDonato, R. Domenech, A.E Duerr, T. E. Katzner, M. J. Lanzone, D.W. LaPlante, C.L. McIntyre, T.A. Miller, R.K. Murphy, A. Shreading, S.J. Slater, J.P. Smith, B.W. Smith, J.W. Watson and B. Woodbridge. 2017. Patterns of spatial distribution of golden eagles across North America: how do they fit into existing landscape-scale mapping patterns? Journal of Raptor Research 51: 197-215.

Schmidt, J.S., C.L. McIntyre, C.A. Roland, M.C. MacCluskie, and M.J. Flamme. 2018. Bottom-up processes drive reproductive success in an apex predator. Ecology and Evolution 2018; 00:1-9.

Schmidt, J.H., E.A. Rextad, C.A. Roland, C.L. McIntyre, M.C. MacCluskie, and M.J. Flamme. 2017. Weather-driven change in primary productivity explains variation in the amplitude of two herbivore population cycles in a boreal system. Oecologia.

Mizel, J. D., J. H. Schmidt, C. L. McIntyre, and M. S. Lindberg. 2017. Subarctic-breeding passerines exhibit phenological resilience to extreme spring conditions. Ecosphere 8(2):e01680. 10.1002/ecs2.1680

McIntyre, C.L. and S.B. Lewis. 2016. Observations of migrating Golden Eagles in eastern interior Alaska offer insights on population size and migration monitoring. Journal of Raptor Research 50: 254-264.

Mizel, J. D., J. H. Schmidt, C. L. Mcintyre, and C. A. Roland. 2016. Rapidly shifting elevational distributions of passerine species parallel vegetation change in the subarctic. Ecosphere 7(3):e01264. 10.1002/ecs2.1264

Doyle, K.M., T.E. Katzner, G.W. Roemer, J.W. Cain, III, B.A. Millsap, C.L. McIntyre, S.A. Sonsthagen, N.B. Fernandez, M. Wheeler, Z. Bulut, P.H. Bloom and J.A. DeWoody. 2016. Genetic structure and viability selection in the golden eagle (Aquila chrysaetos), a vagile raptor with a Holarctic distribution. Conservation Genetics July 2016.

McIntyre, C.L., and M.D. Paulson. 2015. What came first, the nest or the egg? An unusual golden eagle nest observed in Denali National Park and Preserve, Alaska. Journal of Raptor Research 49: 98-101.

Fackler, P.L., K. Pacific, J. Martin, and C. McIntyre. 2014. Efficient use of information in adaptive management with an application to managing recreation near golden eagle nesting sites. PLoS ONE 9(8): e102434. doi:10.1371/journal.pone.0102434.

Schmidt, J.H., C.L. McIntyre, and M.C. MacCluskie. 2013. Accounting for incomplete detection: what are we estimating and how might it affect long-term passerine monitoring programs. Biological Conservation 160: 130-139.

McIntyre, C.L., and J.H. Schmidt. 2012. Ecological and environmental correlates of territory occupancy and breeding performance of migratory Golden Eagles Aquila chrysaetos in interior Alaska. Ibis 154:124-135.

Martin, J., P.L. Fackler, J.D. Nichols, M.C. Runge, C.L. McIntyre, B.L. Lubow, M.C. McCluskie, and J.A. Schmutz. 2011. An adaptive-management framework for optimal control of hiking near Golden Eagle nests in Denali National Park. Conservation Biology 25:316-323.

McIntyre, C.L., D.C. Douglas, and L.G. Adams. 2011. Movements of juvenile Gyrfalcons from western and interior Alaska following departure from their natal areas. Pp. 283-290 in R.T. Watson, T.J. Cade, M. Fuller, G. Hunt, and E. Potapov (eds.), Gyrfalcons and ptarmigan in a changing world. Vol. I. The Peregrine Fund, Boise, ID.

Martin, J., C.L. McIntyre, J.E. Hines, J.D. Nichols, J.A. Schmutz, and M.C. MacCluskie. 2009. Dynamic multistate site occupancy models to evaluate hypotheses relevant to conservation of Golden Eagles in Denali National Park, Alaska. Biological Conservation 142:2726-2731.

McIntyre, C.L., J.M. Withers, and N.J. Gates. 2009. Merlins (Falco columbarius) scavenge road-killed Snowshoe Hares (Lepus americanus) in interior Alaska. Journal of Raptor Research 43:254.

McIntyre, C.L., D.C. Douglas, and L.G. Adams. 2009. Movements of juvenile Gyrfalcons from western and interior Alaska following departure from their natal areas. Journal of Raptor Research 43:99-109.

McIntyre, C.L., D.C. Douglas, and M.W. Collopy. 2008. Movements of Golden Eagles (Aquila chrysaetos) from interior Alaska during their first year of independence. Auk 125:214-224.

McIntyre, C.L., and M.W. Collopy. 2006. Postfledging dependence period of migratory Golden Eagles (Aquila chrysaetos) in Denali National Park and Preserve, Alaska. Auk 123:877-884.

McIntyre, C.L., M.W. Collopy, J.G. Kidd, and A.A. Stickney. 2006. Characteristics of the landscape surrounding Golden Eagle nest sites in Denali National Park and Preserve, Alaska. Journal of Raptor Research 40:46-51.

McIntyre, C.L., M.W. Collopy, and M.S. Lindberg. 2006. Survival probability and mortality of migratory juvenile Golden Eagles from Interior Alaska. Journal of Wildlife Management 70:717-722.

McIntyre, C., K. Steenhof, M.N. Kochert and M.W. Collopy. 2006. Long-term Golden Eagle studies in Denali National Park and Preserve. Alaska Park Science 5:42-45.

Brown, R.J., and C. McIntyre. 2005. New prey species documented for Northern Pike (Esox lucius): Bald Eagle (Haliaeetus leucocephalus). Arctic 58:437.

McCaffery, B.J., and C. McIntyre. 2005. Disparities between results and conclusion: do Golden Eagles warrant special concern based on migration counts in the western United States? Condor 107:469-473.

McIntyre, C.L. 2002. Patterns in nesting area occupancy and reproductive success of Golden Eagles (Aquila chrysaetos) in Denali National Park and Preserve, Alaska, 1988-99. Journal of Raptor Research 36:50-54.

Kochert, M.N., K. Steenhof, C.L. McIntyre, and E.H. Craig. 2002. Golden Eagle (Aquila chrysaetos) in A. Poole, and F. Gill (eds.), The Birds of North America no. 684. The Birds of North America, Inc., Philadelphia, PA.

McIntyre, C.L., and L.G. Adams. 1999. Reproductive characteristics of migratory Golden Eagles in Denali National Park, Alaska. Condor 101:115-123.

McIntyre, C.L., and R.E. Ambrose. 1999. Raptor migration in autumn through the Upper Tanana River Valley, Alaska. Western Birds 30:33-38.

Britten, M.W., C.L. McIntyre, and M. Kralovec. 1995. Satellite radiotelemetry and bird studies in national parks and preserves. Park Science 15:20-24.

Young, D.D., Jr., C.L. McIntyre, P.J. Bente, T.R. McCabe, and R.E. Ambrose. 1995. Nesting by Golden Eagles on the north slope of the Brooks Range in northeastern Alaska. Journal of Field Ornithology 66:373-379.

Swem, T., C. McIntyre, R.J. Ritchie, P.J. Bente, and D.G. Roseneau. 1994. Distribution, abundance, and notes on the breeding biology of Gyrfalcons Falco rusticolus in Alaska. Pp. 437-444 in B.-U. Meyburg and R.D. Chancellor (eds.), Raptor conservation today. World Working Group on Birds of Prey, Berlin, and Pica Press, London.

Denali National Park & Preserve

Last updated: November 12, 2023