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An introduction to the benthic macroinvertebrate community at Appomattox Court House National Historical Park

NPS staff with a net in a stream, stonefly larvae on rock, hellgramite, NPS staff examining net.

The Mid-Atlantic Inventory and Monitoring Network (MIDN) monitors ecosystem health in ten national parks in the Northeast Region. To measure the health of streams and rivers, scientists sample and identify invertebrates living on the bottom of streams. These invertebrates are called “benthic macroinvertebrates.”

Many states and federal agencies use benthic macroinvertebrates to assess stream health and make determinations about whether streams are meeting standards outlined in the federal Clean Water Act. Ultimately, the goal of the benthic macroinvertebrate monitoring program is to provide the best available science so national park managers can better understand and protect park resources.

This brief provides an introduction to benthic macroinvertebrate communities at Appomattox Court House National Historical Park. This information includes a general description of benthic macroinvertebrates and, using data from 2009–2018, a summary of macroinvertebrate community characteristics in the park.

The brief does not address the question of stream ecosystem health — future publications will fill that important role.

So, what are benthic macroinvertebrates?

Blue bar chart showing insects make up 97% of the park's benthic macroinvertebrate community.
Appomattox Court House National Historical Park data (2009-2018)

Benthic macroinvertebrates are invertebrates that live on the bottom of streams and rivers and are visible to the naked eye. They include a large number of taxonomic groups. Macroinvertebrates are useful for ecological assessments because their communities can change dramatically at varying levels of pollution.

Most benthic macroinvertebrates captured in streams at the park are insects. These include the larvae of many common groups like mayflies, caddisflies, stoneflies, and the true flies. Other macroinvertebrates that aren’t insects are also present.

Several different types of macroinvertebrates.






Images not to scale. Photos are just examples of macroinvertebrates and do not necessarily represent specific taxa found in the park. Photos courtesy of J. Gallagher (dragonfly larvae), the Smithsonian Institution (mussel and crayfish), Willamette Biology (worm), and B. Henricks (all others) / Flickr.

What is the most common macroinvertebrate in park streams?

Blue bar chart showing the larvae of "true flies" are the most common macroinvertebrate in park streams.
Appomattox Court House National Historical Park data (2009-2018)

One-half of the benthic macroinvertebrate observations from the park are in the taxonomic order Diptera and are known as “true flies.” True flies include insects like mosquitoes, deer flies, crane flies, and midges. True fly larvae are abundant in streams across the mid-Atlantic region.

Mayflies are also very common at the park. After living up to a year as larvae in a stream, mature mayflies hatch into flying adults. In some species, those adults then complete their life cycle (mate and die) within just a few days.

Which macroinvertebrate group is the most diverse (has the largest # of species or genera)?

Blue bar chart showing that caddisflies are the most diverse group of benthic macroinvertebrates in the park.
Appomattox Court House National Historical Park data (2009-2018)
Taxa richness, or the number of unique species or genera in a sample, is a measure of the community composition of benthic macroinvertebrates. In all, 126 taxa have been identified at the park (including 100 taxa of insects).

The top five most diverse groups (i.e. taxonomic orders) are shown to the right. The top two most diverse groups are caddisflies and true flies. With benthic macroinvertebrates, most individuals that are insects are identified to the genus taxonomic level. Non-insect macroinvertebrates are often not identified below the family level, and therefore, non-insect diversity (not shown) is not as well documented.

What's next with benthic macroinvertebrate monitoring?

Blue line graph showing that relatively few new taxa are being identified with the collection of additional benthic macroinvertebrate samples.
Appomattox Court House National Historical Park data (2009-2018)
The figure to the right shows how the number of discovered insect taxa at the park has increased with each additional macroinvertebrate sample. The number of identified taxa rises quickly in the first 10 samples, then starts to slowly flatten out. This indicates that additional samples are uncovering fewer new taxa.

Benthic macroinvertebrate monitoring efforts continue to focus on measuring the health of stream ecosystems in the park. These data help managers understand the condition and trends of park streams and identify potential stressors to these valuable resources.

Click here for a printable pdf of this article.

For more information about this program, visit the MIDN macroinvertebrate website, read the latest summary report, or contact MIDN Ecologist Nathan Dammeyer.

To learn more about benthic macroinvertebrates, including how to identify them with some stunning images, check out the online Atlas of Common Freshwater Macroinvertebrates of Eastern North America.

Appomattox Court House National Historical Park

Last updated: January 19, 2022