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Fauna Series No. 4


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Cover

Contents

Foreword

Introduction

Population and Mortality

Habits

Food

Elk

Deer

Antelope

Bighorn

Other Larger Mammals

Small Mammals

Birds

Misc. Diet

Conclusions

Bibliography





Fauna of the National Parks — No. 4
Ecology of the Coyote in the Yellowstone
National Park Service Arrowhead


CHAPTER IV:
FOOD


ITEMS IN THE COYOTE DIET

DURING the course of the study 5,086 coyote droppings, containing 8,969 food items, were collected. With the exception of less than 200 of them, containing 185 items, all droppings were gathered from the first of April to about the middle of November. Of the 185 items present in those collected in winter, 119 were deer and elk remains and more than 50 of the remainder were in droppings which had undoubtedly been deposited in the summer and fall. The following table, therefore, except for about 119 items of deer and elk, pictures the food habits during the April-November period.

In addition to the foregoing, more than 200 winter droppings not included in the table on page 43, were examined in the northern part of the park. These contained deer and elk remains. In this area where deer and elk winter, these animals, mainly as carrion, make up practically the entire food supply. In the interior of the park snowshoe hares and mice enter more extensively into the winter diet.

About 3,500 droppings were obtained during 1937, a few in 1936, and the remainder in 1938. The material was not tabulated separately for each year since there was no significant difference in the incidence of the items.

Below are listed the number of individuals of each item found in the droppings. Except in the case of the field mouse and pocket gopher, and in a few instances the ground squirrel and deer mouse, the number of individuals present coincides with the number of droppings in which they occurred. Such items as insects, vegetation, and carrion were arbitrarily listed as number of times occurring, rather than number of individuals, since such material is not otherwise readily comparable with the other food items. For instance, the number of individual grasshoppers, crickets, june beetles, snails, pine nuts, rose seeds, mushrooms, blueberries, and Oregon grape is not given. However, in the discussion of each item the quantity present is usually indicated. The percentages given in the table are based on number of individuals, treating times of occurrence of these few items as individuals. The seats are not always accurately dated. Some collected in spring especially along streams, may be winter droppings. However, the dates of most of them are sufficiently accurate to furnish a picture of coyote diet on a seasonal basis.

Classification of 8,969 individual food items found in 5,086 coyote droppings gathered in Yellowstone National Park

Food items Number of
individuals
Percent
LARGE MAMMALS
Elk, Cervus canadensis nelsoni
Elk Calf
Deer, Odocoileus hemionus macrotis
Deer Fawn
Antelope, Antilocapra americana americana
Antelope Fawn
Mountain Sheep, Ovis canadensis canadensis
Buffalo, Bison bison bison
Moose, Alces americanus shirasi
Domestic cattle
Black Bear, Euarctos americanus cinnamomum
Fragments of large bones
1,153
300
91
2
17
32
2
18
1
5
43
47
12.85
3.34
1.01
.02
.18
.36
.02
.20
.01
.05
.47
.52
SMALL MAMMALS
Field Mouse, Microtus sp
Pocket Gopher, Thomonys fuscus fuscus
Snowshoe Hare, Lepus bairdi bairdi
Marmot, Marmota flaviventris nosophora
Muskrat, Ondatra zibethica osoyoosensis
Ground Squirrel, Citellus armatus richardsoni
Jackrabbit, Lepus townsendii campanius
Porcupine, Erethizon epixanthum epixanthum
Deer Mouse, Peromyscus maniculatus osgoodi
Pine Squirrel, Sciurus hudsonicus ventorum
Beaver, Castor canadensis missouriensis
Coyote, Canis lestes
Cottontail Rabbit, Sylvilagus nuttalli grangeri
Jumping Mouse, Zapus princeps princeps
Chipmunk, Eutamias sp
Woodrat, Neotoma cinerea orolestes
Mink, Mustela vison energumenos
Shrew, Sorex sp
Coney, Ochotona princeps ventorum
Weasel, Mustela frenata ssp
Bat, Myotis sp
House Cat
Flying Squirrel, Glaucomys sabrinus bangsi
Marten (immature), Martes caurina origenes
3,044
1,939
305
120
98
46
37
35
34
25
17
13
10
7
6
4
3
2
1
1
1
1
1
1
33.93
21 61
3.40
1.33
1.09
.51
.41
.39
.37
.27
.18
.14
.11
.07
.06
.04
.03
.02
.01
.01
.01
.01
.01
.01
BIRDS
Duck
Duckling
Bird
Small Bird
Large Bird
Canada Goose, Branta canadensis canadensis
Richardson's Grouse, Dendragapus obscurus richardsoni
Ruffed Grouse, Bonasa umbellus umbelloides
Grouse
Sparrow (immature)
Sparrow
Warbler
Steller's (black-headed) Jay, Cyanocitta stelleri annectens
Spotted sandpiper, Actitis macularia
Short-eared Owl, Asio flammeus flammeus
Eared Grebe, Colymbus nigricollis californicus
Domestic chicken (refuse)
Large bird egg
Small bird egg
Domestic chicken eggshell (refuse)
81
1
62
55
18
13
5
2
5
5
2
2
1
1
2
3
10
7
13
.90
.01
.69
.61
.20
.14
.05
.02
.05
.05
.01
.02
.02
.01
.01
.02
.03
.11
.07
.14
COLD-BLOODED VERTEBRATES
Fish
Garter Snake
12
9
.13
.10
INVERTEBRATES
Grasshopper
Cricket (Anabrus simplex)
June Beetle
Snail
711
123
14
4
7.92
1.37
.15
.04
VEGETABLE MATTER
Grass
Pine Nuts, Pinus albicaulis
Rose Seed (Rosa)
Strawberry
Mushroom
Blueberry, Vaccinium sp
Oregon Grape, Berberis repens
88
51
20
19
4
1
1
.98
.57
.22
.21
.04
.01
.01
MISCELLANEOUS FOOD AND NONFOOD ITEMS
Horse manure
Garbage
Trash
Muskmelon
Apple
Corn (refuse)
Paper
Canvas-leather glove
Rag
Butter wrapper
Twine
Banana peel
Orange peel
Leather (1 piece containing rivet)
Cellophane
Steak bone
Grape seeds
Mouse nest material
7 inches of curtain
Pear
Prune seed
Match
2 sq. inches rubber
Tinfoil
Shoestring
Mud
Paint-covered rag
8 inches of rope
3 sq. inches towel
Lemon rind
Bacon rind
Two pieces of shirt
Canvas
Gunny sack
Isinglass
Botfly larvae
48
12
13
9
7
7
11
6
5
5
4
4
3
3
2
2
2
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
.54
.13
.14
.10
.07
.07
.12
.06
.05
.05
.04
.04
.03
.03
.02
.02
.02
.01
.01
.01
.01
.01
.01
.01
.01
.01
.01
.01
.01
.01
.01
.01
.01
.01
.01
.07







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