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Ecological Restoration

 
Sequoia Seedlings in a Clearing
Sequoia seedlings were planted in clumps to mimic patterns of natural regeneration following fire. © NPS photo.
 
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Restoring Vegetation

Restoration Goals Development in Giant Forest altered the vegetation in several ways. Trees were cleared for buildings and parking lots, leaving distinct openings in the forest canopy. The forest overstory was thinner because trees that threatened human safety and property were removed. Trampling and soil compaction reduced or eliminated the forest understory, including grasses, wildflowers, shrubs, and tree seedlings. The soil seed bank, which influences the regenerative potential of the forest, was likely depleted. Small patches of wetland vegetation were lost where fill was placed over meadow edges or streams.

Fire-caused gaps are colonized by patches of abundant shrub and tree regeneration. © NPs photo by Athena Demetry.
To restore the vegetation, ecologists looked to nature for a disturbance condition that resembled the disturbance caused by human development. In Giant Forest Village, the forest structure was a matrix of mature canopy interspersed with openings, or gaps, where patches of trees were cleared for buildings and parking lots. This condition was similar to surrounding areas of Giant Forest where natural, prescribed fire had killed patches of mature trees, creating openings, or gaps, in the canopy. These fire-caused gaps were colonized by patches of abundant shrub and tree regeneration, particularly giant sequoia, with little regeneration beneath intact canopy. Because gaps caused by fire are of similar size to gaps caused by human development, the vegetation growing within them can be used as a model, or reference condition, for restoration treatments. For treatments involving planting, mimicking the effects of fire is a particularly appropriate model because planting methods --within gaps only, during a short time period, with one to two year-old seedlings-- are similar to the pulse of regeneration occurring within gaps shortly after fire.

Shrub and tree regeneration in fire-caused gaps was mapped with a total station. These patterns of regeneration were used as a model for restoring vegetation in Giant Forest Village. © NPs photo by Athena Demetry.
This reference condition was quantified in 1994 by mapping and measuring woody vegetation in 18 fire-caused gaps in Giant Forest, 7 to 15 years following fire. Gap size was found to account for a significant amount of variability in density, growth rate, and/or cover of pioneer-type tree and shrub species. Learn more from this technical paper (231 KB, PDF format) about the use of fire-caused gaps as a natural disturbance model for restoration.

To summarize, the short-term goal of vegetation restoration in Giant Forest Village is to reproduce the species composition, density, and spatial pattern of regeneration that would result from a natural fire event. The long-term goal is to integrate the site into the natural fire regime typical of surrounding areas of Giant Forest. In other words, the Park is putting in place a structure known to be within the range of natural variability, then allowing natural processes to thin the vegetation. By ensuring a vegetation structure similar to surrounding sites after one fire, the Park maximizes the success of this integration.

Active Restoration or Natural Regeneration?
Because of the duration and severity of changes in developed areas, the Park believed that some degree of human intervention was necessary for the recovery of soils and vegetation in Giant Forest Village. Evidence for this view lies in some formerly developed areas within the grove that were abandoned over 30 years ago and show little natural recovery. However, it was also hypothesized that an acceptable restoration of vegetation might be achieved through less intensive and intrusive means than the seed collection, propagation, planting, seeding, and irrigation process traditionally practiced in the Park’s frontcountry revegetation projects. To address this possibility, an adaptive management approach was taken. The goal of adaptive management was to apply different degrees of active restoration in an experimental manner to determine the minimal intervention necessary to meet the reference condition: natural vegetation in fire-caused gaps.

Three levels of vegetation restoration in Giant Forest Village were tested, in order of increasing human intervention:

1) Restore Soil Only. Actions are limited to regrading to restore natural landforms, amending soils in highly disturbed sites, cultivating, and mulching with litter and duff or wood chips. This is considered the minimal treatment. It was used in four experimental gaps in highly disturbed sites, and also in non-gap areas, former campgrounds abandoned for 30 years or more, and in narrow linear road corridor and trail disturbances through established forest, covering about 42 acres.
Burn piles were placed near mature sequoias to produce heat. The intent was to mimic natural conditions in which heat rises to the canopy, opens the green cones and causes a seed release. © NPs photo by Athena Demetry.
2) Restore Soil and Burn. In addition to actions from treatment (1.), a light fire fuel bed and several large slash piles were imported and burned with the intent of releasing sequoia seed and scarifying the seed bank. This treatment was used in four experimental gaps in highly disturbed sites, covering 2 acres.
3) Restore Soil and Plant. In addition to actions from treatment (1.), active planting occurred. Trees, shrubs, grasses, and wildflowers were propagated from local stock (seed and cuttings collected within Giant Forest) to preserve genetic integrity, and planted in gaps using prescriptions formed from fire-caused reference gaps. Gaps are irrigated for 2 to 3 years to enhance survival. Trees (5 species) were propagated by the USDA-Forest Service in Placerville, California, and planted as 1 or 2 year bare-root or 1-gallon containerized stock. Shrubs (12 species) were propagated by Bitterroot Restoration, Inc. in Corvallis, Montana, and planted as 10 cubic-inch or 1-gallon containerized stock. Grasses and wildflowers (10 species) were propagated by the Natural Resources Conservation Service in Meeker, Colorado, and seeded or planted as plugs. To learn more, view this species list (171 KB, PDF format) of plants used in the restoration. Because this treatment was considered to have the highest probability of success, it was used in the majority of gaps in highly disturbed sites, covering 27 acres.
NPs Natural Resources Personnel quantify vegetation in a gap restored with a burn treatment. © NPs photo by Athena Demetry.
As of December 2001, a total of 7275 trees, 4500 shrubs, and 5395 grasses and wildflowers have been planted, and 22 pounds of seed have been sown. Planting has restored 19 acres of the eventual project total of 27.

The vegetation resulting from these three treatments is being monitored to compare treatment success. First-year monitoring results comparing treatments 2 (burn) and 3 (plant) suggest that both treatments will be successful in producing similar numbers of trees and shrubs as the fire-caused reference gaps 10 years after fire. However, the planting treatment has accelerated the recovery process, with planted gaps having significantly higher plant cover area (that is, the plant canopies are larger) and taller tree seedlings than burned gaps. Learn more from this technical paper (1,131 KB, PDF format) discussing monitoring in detail, including a table of survival rates of planted stock.

Photopoint monitoring of these three treatment sites is ongoing. Four years after restoration, Treatment 1 (soil restoration) continues to show slow vegetative recovery. Treatment 2 (burn) vegetation is showing very rapid growth, perhaps exceeding the planting treatment. Treatment 3 (plant) vegetation is surviving well, but generally shows slower growth than the burn treatment.

 

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