North Cascades National Park: Terrestrial Riparian Arthropod Investigations

Table 2. Pitfall trap captures from nine riparian habitat types in Big Beaver Creek, North Cascades National Park Service Complex, Washington, May - October, 1995.

Species* Habitat Types Species
Totals Family
Totals
AS AT BOG GVL PF SCS SSS TSCS TTF
Amphizoidae (1 species)
Amphizoa insolens LeConte

1

1 1
Anthicidae (3 species)
Anthicus nanus LeConte

19
1

20
Eurygenius campanulatus LeConte

373
1

373

Ischalia vancouverensis Harrington 1

1 2 395
Buprestidae (1 species)
Agrilus politus (Say)

1

1 1
Byrrhidae (7 species)
Byrrhus kirbyi LeConte 1

1

Curimopsis albonotata LeConte

1

1

Cytilus alternatus Say

8 36

44

Exomella pleuralis (Casey)

1
1

Listemus acuminatus (Mannerheim) 1 1

2

Morychus aeneolus LeConte

3

3

Morychus oblongus LeConte

1

1 53
Cantharidae (4 species)
Cantharis oregonus LeConte

1

1

Malthodes sp.

1

1

Podabrus conspiratus Fall

1

1

Podabrus piniphilus Dejean
4

3

2 9 12
Carabidae (54 species)
Agonum affine Kirby

1

1

Agonum brevicolle Dejean

10

138 57

205

Agonum ferruginosum Dejean 1

17 13 4
35

Agonum piceolum LeConte 2 1

3

Agonum thoreyi Dejean

1 17

18

Amara littoralis Mannerheim

2

2

Amara sanjuanensis Hatch
1

1

Anchomenus quadratus (LeConte)

1

1

Anisodactylus binotatus Fabricius

1

1

Apristus constrictus Casey

1

1

Bembidion breve (Motschulsky)

1

1

Bembidion concretum Casey

1

1

Bembidion convexulum Hayward

3
1
4

Bembidion fortestriatum Motschulsky

9

5 11 5
30

Bembidion hesperum Casey

1

1

Bembidion inaequale Say

1

1

Bembidion incrematum LeConte

6 11

17

Bembidion iridescens LeConte
3

3

Bembidion kuprianovi Mannerheim 30

2

3 35

Bembidion semipunctatum Kirby

1

1

Bembidion planatum LeConte

40

40

Bembidion planiusculum Mannerheim

2

2

Bembidion quadrifoveolatum Mannerheim

1

1

Bembidion quadrulum LeConte

1

1

Blethisa oregonensis LeConte

65

65

Bradycellus conformis Fall

1

8
10
19

Bradycellus lecontei Csiki

2

2

Bradycellus nigrinus Dejean

26 15

41

Chlaenius interruptus Horn

3

21 12

36

Cicindela depressula Casey

26

26

Cicindela oregona LeConte

23

23

Diplous atterrimus Dejean

4

4

Elaphrus clairvillei Kirby

2

1 1

4

Elaphrus purpurans Hausen 1

1

Harpalus cordifer Notman 6

6

Harpalus somnulentus Dejean

1
1
2

Leistus ferruginosus Mannerheim 12 3

1

3 22 41

Loricera decempunctata Eschscholtz 3 6

18 31 23 1 82

Nebria mannerheimi Fischer

40

40

Nebria sahlbergi Fischer

20

20

Notiophilus sylvaticus Eschscholtz

1

2 3

Opisthius richardsoni Kirby

14

14

Patrobus fossifrons dimorphicus Darl. 2

1 3

Pterostichus adstrictus Eschscholtz 21

21 2
2 46

Pterostichus herculaneus Mannerheim 9 6 1 2 27

4 18 67

Pterostichus neobrunneus Lindroth 2 1

53

62 128

Pterostichus riparius Dejean 28 1
4
13
23 5 74

Scaphinotus angulatus Harris
7

2

9

Scaphinotus angusticollis Mannerheim 35 86

343
1
211 676

Scaphinotus marginatus Fischer 30 8
1 3 9 7 1 17 76

Synuchus impunctatus Say

1

2 3

Trechus chalybeus Dejean 4 6

18 6 38
72

Trechus oregonensis Hatch 5 1

1 53
60
Trichocellus cognatus Gyllenhal

9

9 2058
Cerambycidae (5 species)
Brachyleptura dehiscens (LeConte)

1

1

Leptura obliterata Haldeman
1

1

Plectura spinicauda Mannerheim
3

3

Xestoleptura crassipes (LeConte)

1

1
1
3

Xestoleptura tibialis LeConte

1

1 9
Chrysomelidae (10 species)
Altica corni Woods
1

1

Altica tombacina Mannerheim

1 1
2

4

Chaetocnema irregularis LeConte

1

1

2

Chrysomela mainensis Bechyne 1

1

Crepidodera nana (Say)

1

1

Hippuriphila mancula LeConte

2

2

Macrohaltica caurina Blake

2

2

Plateumaris nitida Germar

1

1

Pyrrhalta punctipennis Mannerheim,<
>aberration pallida Beller & Hatch

3

3

Pyrrhalta spiraeophila Hatch & Beller

1

1 18
Ciidae (1 species)
Cis maritimus (Hatch)

1 1 1
Coccinellidae (3 species)
Hippodamia washingtoni Timberlake

2

2

Scymnus caurinus Horn

2

2

Stethorus punctum picipes Casey 1

1 5
Cryptophagidae (11 species)
Anchicera ephippiata Zimmerman

3
3

Anchicera kamtschatica Motschulsky 6

12
18

Anchicera ochracea Zimmerman

1

1
2

Anchicera postpallens Casey

12 1

13

Antherophagus ochraceus Melsh.

1

1

Atomaria constricta Casey

1

1

Atomaria quadricollis Casey

1
1

Cryptophagus confertus Casey 1 3

1

5 10

Cryptophagus lapponicus Gyllenhal
1

2 3

Cryptophagus tuberculosus Maklin 1

1

Henotiderus lorna Hatch

2

2 55
Curculionidae (8 species)
Baris sparsa LeConte

1

1

Cryptorhynchus lapathi Linnaeus

1 5
6

Geoderces horni Van Dyke

2

1 3

Lepesoma lecontei Casey

2

2

Lepesoma verrucifera Casey

2 2

Rhyncolus brunneus Mannerheim 1 1

3
1
6 12

Steremnius carinatus Boheman 1 4

18
1
7 31

Sthereus horridus (Mannerheim) 22 11

2

5 40 97
Dytiscidae (7 species)
Agabus austinii Sharp

1
1

Agabus strigulosus (Crotch)

1
3
4

Agabus sp. (female) 1

1

2

Dytiscus sp.

1

1

Graphoderus perplexus Sharp

1

1

Hydroporus pacificus Fall

1
1

Hydroporus sp.

1

1 11
Elateridae (24 species)
Agriotes ferrugineipennis LeConte
1

1

Ampedus rhodopus LeConte

1

1

Athous rufiventris Eschscholtz

1

1
2

Athous vittiger LeConte 1

1 2

Cardiophorus amplicollis Motschulsky

10

10

Cardiophorus propinquus Hatch

246

246

Ctenicera aeripennis (Hatch)

1

1

Ctenicera angusticollis Mannerheim 5

1

1 7

Ctenicera opacula (LeConte)
1

1

Ctenicera volitans Eschscholtz
2

2

Dalopius maritimus Brown
2
1

3

Eanus striatipennis Brown

4

4

Hemicrepidius pallidipennis Mannerheim 3

3

Hypnoidus bicolor Eschscholtz

6

6

Hypolithus dispersus Horn
1
88

89

Hypolithus musculus Eschscholtz

54

54

Hypolithus nocturnus Eschscholtz

1

1

Hypolithus squalidus LeConte

3

3

Hypolithus sp.

31

31

Ligmargus funebris Candeze

125

125

Megapenthes caprella (LeConte)

1

1

Migiwa striatulus (LeConte)

9

9

Negastrius ornatus (LeConte)

20

20

Zorochrus caurinus Horn

6

1

7 629
Erotylidae (1 species)
Triplax antica LeConte 1

1 1
Histeridae (1 species)
Hypocaccus bigemmeus LeConte

2

2 2
Hydraenidae (2 species)
Hydraena vandykei vandykei d'Orchymont

2 5

7

Ochthebius cribricollis LeConte

1

1 8
Hydrophilidae (4 species)
Cercyon adumbratum Mannerheim 3

3

Crenitis paradigma d'Orchymont

1
4
5

Cymbiodyta acuminata Fall

1

1

Helophorus auricollis Eschscholtz

2

2 11
Lampyridae (1 species)
Phausis skelleyi Fender
2

2 2
Latridiidae (5 species)
Enicmus cordatus Belon

2

1 3

Melanopthalma americana Mannerheim

11

45 6 5
67

Melanopthalma distinguenda Comolli

8

1 1

10

Melanopthalma gibbosa Herbst

2

1
1
4

Steposthethus liratus LeConte 1

1 85
Leiodidae (20 species)
Agathidium californicum Horn 1

1 2

Agathidium concinnum Mannerheim 1

1 2

Agathidium contiguum Fall
1

1

Agathidium jasperinum Fall
3 1
1 1

6

Anisotoma confusa Horn

1

1

Catops basilaris Say 14

14

Catops egenus Horn 36 5

26 22
89

Catops simplex Say 1

1 2
4

Catops spp. 882 2088 84 57 203 74 328 273 540 4529

Catoptrichus frankenhaeuseri (Mannerheim) 2 18

1

21

Colon inerme Mannerheim
4

2

6

Colon magnicolle Mannerheim
2
1
1 2 3
9

Colon nevadense/I> Horn
3

3

Colon serripoides Hatch

4 3 5
12

Colon sp.
1

2

2 5

Leiodes alesi Baronowski
1

1

Leiodes cascadensis Baronowski 8

2 3 7 2 22

Leiodes lateritia (Mannerheim) 1

2 3

Leiodes puncticollis (Thomson)
2

3

1 6

Leptinus occidentamericanus Peck 4 1 2 1

1 9

Platycholeus opacellus Fall

1

1 4746
Lycidae (1 species)
Dictyopterus simplicipes Mannerheim 2

2 2
Melandryidae (1 species)
Xylita laevigata Hellenius

1

1 1
Mordellidae (1 species)
Mordella atrata Melsheimer

1

1 1
Nitidulidae (1 species)
Epuraea avara Randall

1

1 1
Oedemeridae (4 species)
Calopus angustus LeConte

1 1

Ditylus gracilis LeConte 2 5

1

1 3 12

Ditylus quadricollis LeConte 7 6
19

1 2
35

Xanthochroa testacea Horn

` 1 49
Phalacridae (2 species)
Phalacrus pencillatus Say

2

2

Stilbus apicalis Melsheimer
1 1

3

5 7
Ptiliidae (4 species)
Acrotrichis cognata Matthews
10 7
1
1

19

Acrotrichis henrici Matthews

1

9
10

Acrotrichis vicina Matthews 8 8 1

5 6 4 1 33

Ptenidium pusillum Gyllenhal

1

1 5 7
14 76
Ptilodactylidae (1 species)
Araeopidius monachus LeConte

1

1 1
Pyrochroidae (2 species)
Dendroides ephemeroides Mannerheim 2 2
1

5

Pedilus jonae Young 1

1 6
Pythidae (1 species)
Priognathus monilicornis Randall

1 1 1
Scarabaeidae (4 species)
Aegialia lacustris LeConte

2

2

Aegialia opaca Brown

1

1

Aphodius opacus LeConte
2

1

1 4

Onthophagus nuchicornis Linnaeus

1

1 8
Scirtidae (3 species)
Cyphon brevicollis LeConte 1 1

1 3

6

Cyphon padi Linnaeus

1

1

Cyphon variabilis Thunberg

5

1
2
8 15
Scolytidae (1 species)
Gnathotrichus retusus (LeConte)

1

1 1
Scydmaenidae (3 species)
Scydmaenus californicus Motschulsky 4 7
1 2

1 1 16

Scydmaenus fuchsi Brendel

1 1

Veraphis mirabilis Marsh 2 15

4

21 38
Silphidae (3 species)
Nicrophorus defodiens Mannerheim 40 51 57 5 10 94 96 26 22 401

Nicrophorus investigator Zetterstedt 5 1 17 6
6 7
2 44

Nicrophorus spp. (in alcohol)

75
41 336 140 989 90 780

Thanatophilus lapponicus (Herbst)

2

2 1227
Sphaeritidae (1 species)
Sphaerites politus Mannerheim 2

2 2
Staphylinidae (97 species)
Acidota crenata Fabricius 7 1

1

1 10

Actium barri Park & Wagner
1

3

3 7

Aleochara bilineata Gyllenhal

2
1

3

Aleochara bimaculata Gravenhorst

1 1

2

Aleocharinae 348 616 18 7 138 47 99 117 117 1507

Amphicroum maculatum Horn

1 1

Anthobium clarkae Hatch 2

5

1 8

Anthobium reflexicolle Casey 14

1

11 26

Anthobium sinuosum Hatch 1 1

2

Baeocera humeralis Fall 1
2

3

Batrisodes albionicus (Aube)
1

3
4

Bisnius siegwaldi (Mannerheim) 14 1

1 1 2
5 24

Bledius suturalis LeConte

6

6

Bolitobius kremeri Maklin 2

1

2 5

Bryophacis discalis (Hatch)
1

1

Bryophacis punctatissimus Hatch 2

1

3

Bryophacis punctulatus Hatch 3

3

Cupila excavata Park & Wagner 2
4
2 2
1
11

Cypha crotchi Horn

2

2

Elonium NEAR barri (Hatch) 2

7 9

Elonium rugosa (Hatch)

1

1

Empelus brunnipennis Mannerheim 2

1 1

3 7

Erichsonius cinerascens Gravenhorst

10

1 3

14

Eusphalerum fenyesi Bernh. 2 1

3

Eusphalerum pothos Mannerheim 2 1 2

5

Gabrius cushmani Hatch 2 25

1

2 6 36

Gabrius picipennis Maklin 2
5

1 1 15
24

Gabrius seattlensis Hatch 41

2 1 4
48

Gabrius shulli Hatch
9

3 12

Hemiquedius fusculus (LeConte)

2

2

Ischnosoma fimbriatum Campbell 1 1

1

3

Ischnosoma pictum (Horn) 2

1

1
4

Ischnosoma splendidus (Gravenhorst)

4
4

Lathrobium punctulatum? LeConte

3

3

Lathrobium vancouveri Casey
1

1

Lithocaris capitula Casey

1

1

2

Lobrathium sp.

1

1

Lordithon fungicola Campbell 39 2
4 13
1 11 7 77

Lordithon poecilus Mannerheim 13 7

3
1
7 31

Lordithon thoracicus Fabricus 1

1

2

Lucifotychus cognatus LeConte 3

1
1
1 4 10

Lucifotychus impellus Park & Wagner

1

1

Mathrilaeum subcostatum Maklin
1

1
1
1 4

Megarthrus pictus Motschulsky 7

9 16

Megarthrus sinuaticollis Boisduval & Lacordaire 26
2
3

1 32

Microedus austinianus LeConte

1

1

Micropeplus minor Campbell

1
1

Micropeplus nelsoni Campbell 4

4

Mycetoporus americanus Erichson 2

2

Mycetoporus bipunctatus Campbell

1 1

Mycetoporus maculicollis LeConte

1

1

Mycetoporus pacificus Campbell 3 1

2 6

Olophrum consimile Gyllenhal

6

6

Omalium foraminosum Maklin 3 1

4

Ontholestes cingulatus Gravenhorst

1

1 2

Oropodes dybasi Grigarick & Schuster

1

1

Oropus striatus (LeConte) 2 11

3

1 4 21

Orus punctatus Casey

1

1

Oxyporus occipitalis Fauvel

1 1

Oxytelus laqueatus Marsham 37 5

1
2 1 46

Pelecomalium testaceum Mannerheim 2

2

Philonthus crotchi Horn

15

9 6 5 1 36

Philonthus cruentatus (Gmelin)

2

2

Philonthus duplicatus Bernhauer & Schubert

1
1

Philonthus furvus Nordmann 1

1
1
3

Philonthus varians Paykull
1

1

Proteinus basalis Maklin 12 8

2
1 1 4 28

Proteinus collaris Hatch 63 82

4 26 175

Proteinus limbatus Maklin 7 1

1 11 20

Pseudopsis sulcata Newman 2

2

Quedius aenescens Maklin 1

1

Quedius breviceps? Casey 1

1

Quedius crescenti Hatch 1

1 2

Quedius fulvicollis (Stephens)

10
10

Quedius griffinae Hatch 1

1 2

Quedius horni Hatch 21 11
2
2 1 15 7 59

Quedius nevadensis Casey

1

1

Quedius oculeus Casey

1

1

Reichenbachia albionica Motschulsky 17
4 1
35 44 68 2 171

Sonoma hespera Park & Wagner

1
1

2

Staphylinus pleuralis LeConte 5 16

28

2 31 82

Staphylinus rutilicauda Horn 1

1

2

Stenus juno Fabricius

6

6

Stenus laccophilus Casey 1

9 3 7
20

Stenus occidentalis Casey

10

8

18

Stenus subgriseus Casey

5

23 3

31

Subhaida ingrata (Hatch)
1

1

Tachinus basalis Erichson 3 1
1 1

3 9

Tachinus crotchii Horn 61 50 `0 6 `2 8 `1 9 12 181

Tachinus maculicollis Maklin 8 1

1 7 17

Tachinus nigricornis Mannerheim

2

6 8

Tachinus semirufus Horn

10

7 17

Tachyporus canadensis Campbell

2 7 6
15

Tachyporus chrysomelinus Linnaeus

1
1

Tachyporus maculicollis Campbell

1
1
2

Tachyporus mexicanus Sharp

2 3

5

Trichophya pilicornis Gyllenhal

5
1

6

Unamis fulvipes Fall

1

1 3020
Tenebrionidae (2 species)
Helops pernitens LeConte

2 2

Scaphidema pictum LeConte

13

13 15
Throscidae (2 species)
Aulonthroscus validus LeConte

1

1

Pactopus hornii LeConte

2

2 3
Trogositidae (1 species)
Temnochila chlorodia Mannerheim

1

1 1
Zopheridae (1 species)
Phellopsis porcata LeConte

3

3 3

* Catops spp., Nicrophorus spp., and unidentified Aleocharinae were excluded from the species counts.

http://www.nps.gov/noca/arthropod2-t2.htm
Last Updated: 25-Nov-1998

Species*	Habitat Types									Species Totals	Family Totals
Species*	AS	AT	BOG	GVL	PF	SCS	SSS	TSCS	TTF	Species Totals	Family Totals
Amphizoidae (1 species)
Amphizoa insolens LeConte				1						1	1
Anthicidae (3 species)
Anthicus nanus LeConte				19		1				20
Eurygenius campanulatus LeConte				373		1				373
Ischalia vancouverensis Harrington	1								1	2	395
Buprestidae (1 species)
Agrilus politus (Say)			1							1	1
Byrrhidae (7 species)
Byrrhus kirbyi LeConte	1									1
Curimopsis albonotata LeConte				1						1
Cytilus alternatus Say						8	36			44
Exomella pleuralis (Casey)								1		1
Listemus acuminatus (Mannerheim)	1	1								2
Morychus aeneolus LeConte				3						3
Morychus oblongus LeConte				1						1	53
Cantharidae (4 species)
Cantharis oregonus LeConte				1						1
Malthodes sp.				1						1
Podabrus conspiratus Fall					1					1
Podabrus piniphilus Dejean		4			3				2	9	12
Carabidae (54 species)
Agonum affine Kirby							1			1
Agonum brevicolle Dejean			10			138	57			205
Agonum ferruginosum Dejean	1					17	13	4		35
Agonum piceolum LeConte	2	1								3
Agonum thoreyi Dejean						1	17			18
Amara littoralis Mannerheim						2				2
Amara sanjuanensis Hatch		1								1
Anchomenus quadratus (LeConte)					1					1
Anisodactylus binotatus Fabricius					1					1
Apristus constrictus Casey			1							1
Bembidion breve (Motschulsky)			1							1
Bembidion concretum Casey						1				1
Bembidion convexulum Hayward						3		1		4
Bembidion fortestriatum Motschulsky			9			5	11	5		30
Bembidion hesperum Casey				1						1
Bembidion inaequale Say				1						1
Bembidion incrematum LeConte						6	11			17
Bembidion iridescens LeConte		3								3
Bembidion kuprianovi Mannerheim	30			2					3	35
Bembidion semipunctatum Kirby							1			1
Bembidion planatum LeConte				40						40
Bembidion planiusculum Mannerheim				2						2
Bembidion quadrifoveolatum Mannerheim							1			1
Bembidion quadrulum LeConte				1						1
Blethisa oregonensis LeConte						65				65
Bradycellus conformis Fall			1			8		10		19
Bradycellus lecontei Csiki							2			2
Bradycellus nigrinus Dejean						26	15			41
Chlaenius interruptus Horn			3			21	12			36
Cicindela depressula Casey				26						26
Cicindela oregona LeConte				23						23
Diplous atterrimus Dejean				4						4
Elaphrus clairvillei Kirby			2			1	1			4
Elaphrus purpurans Hausen	1									1
Harpalus cordifer Notman	6									6
Harpalus somnulentus Dejean						1		1		2
Leistus ferruginosus Mannerheim	12	3			1			3	22	41
Loricera decempunctata Eschscholtz	3	6				18	31	23	1	82
Nebria mannerheimi Fischer					40					40
Nebria sahlbergi Fischer					20					20
Notiophilus sylvaticus Eschscholtz					1				2	3
Opisthius richardsoni Kirby				14						14
Patrobus fossifrons dimorphicus Darl.	2								1	3
Pterostichus adstrictus Eschscholtz	21					21	2		2	46
Pterostichus herculaneus Mannerheim	9	6	1	2	27			4	18	67
Pterostichus neobrunneus Lindroth	2	1			53				62	128
Pterostichus riparius Dejean	28	1		4		13		23	5	74
Scaphinotus angulatus Harris		7			2					9
Scaphinotus angusticollis Mannerheim	35	86			343		1		211	676
Scaphinotus marginatus Fischer	30	8		1	3	9	7	1	17	76
Synuchus impunctatus Say					1				2	3
Trechus chalybeus Dejean	4	6				18	6	38		72
Trechus oregonensis Hatch	5	1						1	53		60
Trichocellus cognatus Gyllenhal						9				9	2058
Cerambycidae (5 species)
Brachyleptura dehiscens (LeConte)				1						1
Leptura obliterata Haldeman		1								1
Plectura spinicauda Mannerheim		3								3
Xestoleptura crassipes (LeConte)			1			1		1		3
Xestoleptura tibialis LeConte			1							1	9
Chrysomelidae (10 species)
Altica corni Woods		1								1
Altica tombacina Mannerheim			1	1		2				4
Chaetocnema irregularis LeConte			1			1				2
Chrysomela mainensis Bechyne	1									1
Crepidodera nana (Say)						1				1
Hippuriphila mancula LeConte						2				2
Macrohaltica caurina Blake				2						2
Plateumaris nitida Germar			1							1
Pyrrhalta punctipennis Mannerheim,< >aberration pallida Beller & Hatch							3			3
Pyrrhalta spiraeophila Hatch & Beller			1							1	18
Ciidae (1 species)
Cis maritimus (Hatch)									1	1	1
Coccinellidae (3 species)
Hippodamia washingtoni Timberlake							2			2
Scymnus caurinus Horn					2					2
Stethorus punctum picipes Casey	1									1	5
Cryptophagidae (11 species)
Anchicera ephippiata Zimmerman								3		3
Anchicera kamtschatica Motschulsky	6							12		18
Anchicera ochracea Zimmerman			1					1		2
Anchicera postpallens Casey						12	1			13
Antherophagus ochraceus Melsh.				1						1
Atomaria constricta Casey				1						1
Atomaria quadricollis Casey								1		1
Cryptophagus confertus Casey	1	3			1				5	10
Cryptophagus lapponicus Gyllenhal		1							2	3
Cryptophagus tuberculosus Maklin	1									1
Henotiderus lorna Hatch						2				2	55
Curculionidae (8 species)
Baris sparsa LeConte				1						1
Cryptorhynchus lapathi Linnaeus							1	5		6
Geoderces horni Van Dyke					2				1	3
Lepesoma lecontei Casey					2					2
Lepesoma verrucifera Casey									2	2
Rhyncolus brunneus Mannerheim	1	1			3		1		6	12
Steremnius carinatus Boheman	1	4			18		1		7	31
Sthereus horridus (Mannerheim)	22	11			2				5	40	97
Dytiscidae (7 species)
Agabus austinii Sharp								1		1
Agabus strigulosus (Crotch)						1		3		4
Agabus sp. (female)	1						1			2
Dytiscus sp.						1				1
Graphoderus perplexus Sharp			1							1
Hydroporus pacificus Fall								1		1
Hydroporus sp.				1						1	11
Elateridae (24 species)
Agriotes ferrugineipennis LeConte		1								1
Ampedus rhodopus LeConte				1						1
Athous rufiventris Eschscholtz				1				1		2
Athous vittiger LeConte	1								1	2
Cardiophorus amplicollis Motschulsky				10						10
Cardiophorus propinquus Hatch				246						246
Ctenicera aeripennis (Hatch)							1			1
Ctenicera angusticollis Mannerheim	5				1				1	7
Ctenicera opacula (LeConte)		1								1
Ctenicera volitans Eschscholtz		2								2
Dalopius maritimus Brown		2		1						3
Eanus striatipennis Brown			4							4
Hemicrepidius pallidipennis Mannerheim	3									3
Hypnoidus bicolor Eschscholtz				6						6
Hypolithus dispersus Horn		1		88						89
Hypolithus musculus Eschscholtz				54						54
Hypolithus nocturnus Eschscholtz				1						1
Hypolithus squalidus LeConte				3						3
Hypolithus sp.					31					31
Ligmargus funebris Candeze				125						125
Megapenthes caprella (LeConte)			1							1
Migiwa striatulus (LeConte)				9						9
Negastrius ornatus (LeConte)				20						20
Zorochrus caurinus Horn				6			1			7	629
Erotylidae (1 species)
Triplax antica LeConte	1									1	1
Histeridae (1 species)
Hypocaccus bigemmeus LeConte				2						2	2
Hydraenidae (2 species)
Hydraena vandykei vandykei d'Orchymont						2	5			7
Ochthebius cribricollis LeConte						1				1	8
Hydrophilidae (4 species)
Cercyon adumbratum Mannerheim	3									3
Crenitis paradigma d'Orchymont						1		4		5
Cymbiodyta acuminata Fall							1			1
Helophorus auricollis Eschscholtz						2				2	11
Lampyridae (1 species)
Phausis skelleyi Fender		2								2	2
Latridiidae (5 species)
Enicmus cordatus Belon					2				1	3
Melanopthalma americana Mannerheim			11			45	6	5		67
Melanopthalma distinguenda Comolli			8			1	1			10
Melanopthalma gibbosa Herbst			2			1		1		4
Steposthethus liratus LeConte	1									1	85
Leiodidae (20 species)
Agathidium californicum Horn	1								1	2
Agathidium concinnum Mannerheim	1								1	2
Agathidium contiguum Fall		1								1
Agathidium jasperinum Fall		3	1		1	1				6
Anisotoma confusa Horn					1					1
Catops basilaris Say	14									14
Catops egenus Horn	36	5					26	22		89
Catops simplex Say	1						1	2		4
Catops spp.	882	2088	84	57	203	74	328	273	540	4529
Catoptrichus frankenhaeuseri (Mannerheim)	2	18			1					21
Colon inerme Mannerheim		4					2			6
Colon magnicolle Mannerheim		2		1		1	2	3		9
Colon nevadense/I> Horn		3								3
Colon serripoides Hatch						4	3	5		12
Colon sp.		1			2				2	5
Leiodes alesi Baronowski		1								1
Leiodes cascadensis Baronowski	8					2	3	7	2	22
Leiodes lateritia (Mannerheim)	1								2	3
Leiodes puncticollis (Thomson)		2			3				1	6
Leptinus occidentamericanus Peck	4	1	2	1					1	9
Platycholeus opacellus Fall				1						1	4746
Lycidae (1 species)
Dictyopterus simplicipes Mannerheim	2									2	2
Melandryidae (1 species)
Xylita laevigata Hellenius						1				1	1
Mordellidae (1 species)
Mordella atrata Melsheimer			1							1	1
Nitidulidae (1 species)
Epuraea avara Randall			1							1	1
Oedemeridae (4 species)
Calopus angustus LeConte									1	1
Ditylus gracilis LeConte	2	5			1			1	3	12
Ditylus quadricollis LeConte	7	6		19			1	2		35
Xanthochroa testacea Horn									`	1	49
Phalacridae (2 species)
Phalacrus pencillatus Say						2				2
Stilbus apicalis Melsheimer		1	1				3			5	7
Ptiliidae (4 species)
Acrotrichis cognata Matthews		10	7		1		1			19
Acrotrichis henrici Matthews			1					9		10
Acrotrichis vicina Matthews	8	8	1			5	6	4	1	33
Ptenidium pusillum Gyllenhal			1			1	5	7		14	76
Ptilodactylidae (1 species)
Araeopidius monachus LeConte			1							1	1
Pyrochroidae (2 species)
Dendroides ephemeroides Mannerheim	2	2		1						5
Pedilus jonae Young	1									1	6
Pythidae (1 species)
Priognathus monilicornis Randall									1	1	1
Scarabaeidae (4 species)
Aegialia lacustris LeConte				2						2
Aegialia opaca Brown				1						1
Aphodius opacus LeConte		2			1				1	4
Onthophagus nuchicornis Linnaeus						1				1	8
Scirtidae (3 species)
Cyphon brevicollis LeConte	1	1				1	3			6
Cyphon padi Linnaeus			1							1
Cyphon variabilis Thunberg			5			1		2		8	15
Scolytidae (1 species)
Gnathotrichus retusus (LeConte)						1				1	1
Scydmaenidae (3 species)
Scydmaenus californicus Motschulsky	4	7		1	2			1	1	16
Scydmaenus fuchsi Brendel									1	1
Veraphis mirabilis Marsh	2	15			4					21	38
Silphidae (3 species)
Nicrophorus defodiens Mannerheim	40	51	57	5	10	94	96	26	22	401
Nicrophorus investigator Zetterstedt	5	1	17	6		6	7		2	44
Nicrophorus spp. (in alcohol)			75		41	336	140	989	90	780
Thanatophilus lapponicus (Herbst)						2				2	1227
Sphaeritidae (1 species)
Sphaerites politus Mannerheim	2									2	2
Staphylinidae (97 species)
Acidota crenata Fabricius	7	1			1				1	10
Actium barri Park & Wagner		1			3				3	7
Aleochara bilineata Gyllenhal				2		1				3
Aleochara bimaculata Gravenhorst			1	1						2
Aleocharinae	348	616	18	7	138	47	99	117	117	1507
Amphicroum maculatum Horn									1	1
Anthobium clarkae Hatch	2				5				1	8
Anthobium reflexicolle Casey	14			1					11	26
Anthobium sinuosum Hatch	1	1								2
Baeocera humeralis Fall	1		2							3
Batrisodes albionicus (Aube)		1						3		4
Bisnius siegwaldi (Mannerheim)	14	1			1	1	2		5	24
Bledius suturalis LeConte				6						6
Bolitobius kremeri Maklin	2				1				2	5
Bryophacis discalis (Hatch)		1								1
Bryophacis punctatissimus Hatch	2				1					3
Bryophacis punctulatus Hatch	3									3
Cupila excavata Park & Wagner	2		4		2	2		1		11
Cypha crotchi Horn				2						2
Elonium NEAR barri (Hatch)	2								7	9
Elonium rugosa (Hatch)			1							1
Empelus brunnipennis Mannerheim	2			1	1				3	7
Erichsonius cinerascens Gravenhorst			10			1	3			14
Eusphalerum fenyesi Bernh.	2	1								3
Eusphalerum pothos Mannerheim	2	1	2							5
Gabrius cushmani Hatch	2	25			1			2	6	36
Gabrius picipennis Maklin	2		5			1	1	15		24
Gabrius seattlensis Hatch	41					2	1	4		48
Gabrius shulli Hatch		9							3	12
Hemiquedius fusculus (LeConte)				2						2
Ischnosoma fimbriatum Campbell	1	1					1			3
Ischnosoma pictum (Horn)	2				1			1		4
Ischnosoma splendidus (Gravenhorst)								4		4
Lathrobium punctulatum? LeConte						3				3
Lathrobium vancouveri Casey		1								1
Lithocaris capitula Casey			1				1			2
Lobrathium sp.					1					1
Lordithon fungicola Campbell	39	2		4	13		1	11	7	77
Lordithon poecilus Mannerheim	13	7			3		1		7	31
Lordithon thoracicus Fabricus	1						1			2
Lucifotychus cognatus LeConte	3			1		1		1	4	10
Lucifotychus impellus Park & Wagner				1						1
Mathrilaeum subcostatum Maklin		1			1		1		1	4
Megarthrus pictus Motschulsky	7								9	16
Megarthrus sinuaticollis Boisduval & Lacordaire	26		2		3				1	32
Microedus austinianus LeConte				1						1
Micropeplus minor Campbell								1		1
Micropeplus nelsoni Campbell	4									4
Mycetoporus americanus Erichson	2									2
Mycetoporus bipunctatus Campbell									1	1
Mycetoporus maculicollis LeConte							1			1
Mycetoporus pacificus Campbell	3	1							2	6
Olophrum consimile Gyllenhal							6			6
Omalium foraminosum Maklin	3	1								4
Ontholestes cingulatus Gravenhorst			1						1	2
Oropodes dybasi Grigarick & Schuster				1						1
Oropus striatus (LeConte)	2	11			3			1	4	21
Orus punctatus Casey			1							1
Oxyporus occipitalis Fauvel									1	1
Oxytelus laqueatus Marsham	37	5				1		2	1	46
Pelecomalium testaceum Mannerheim	2									2
Philonthus crotchi Horn			15			9	6	5	1	36
Philonthus cruentatus (Gmelin)			2							2
Philonthus duplicatus Bernhauer & Schubert								1		1
Philonthus furvus Nordmann	1					1		1		3
Philonthus varians Paykull		1								1
Proteinus basalis Maklin	12	8			2		1	1	4	28
Proteinus collaris Hatch	63	82						4	26	175
Proteinus limbatus Maklin	7	1						1	11	20
Pseudopsis sulcata Newman	2									2
Quedius aenescens Maklin	1									1
Quedius breviceps? Casey	1									1
Quedius crescenti Hatch	1								1	2
Quedius fulvicollis (Stephens)								10		10
Quedius griffinae Hatch	1								1	2
Quedius horni Hatch	21	11		2		2	1	15	7	59
Quedius nevadensis Casey				1						1
Quedius oculeus Casey					1					1
Reichenbachia albionica Motschulsky	17		4	1		35	44	68	2	171
Sonoma hespera Park & Wagner			1		1					2
Staphylinus pleuralis LeConte	5	16			28			2	31	82
Staphylinus rutilicauda Horn	1			1						2
Stenus juno Fabricius						6				6
Stenus laccophilus Casey	1					9	3	7		20
Stenus occidentalis Casey			10				8			18
Stenus subgriseus Casey			5			23	3			31
Subhaida ingrata (Hatch)		1								1
Tachinus basalis Erichson	3	1		1	1				3	9
Tachinus crotchii Horn	61	50	`0	6	`2	8	`1	9	12	181
Tachinus maculicollis Maklin	8	1						1	7	17
Tachinus nigricornis Mannerheim					2				6	8
Tachinus semirufus Horn					10				7	17
Tachyporus canadensis Campbell						2	7	6		15
Tachyporus chrysomelinus Linnaeus								1		1
Tachyporus maculicollis Campbell						1		1		2
Tachyporus mexicanus Sharp						2	3			5
Trichophya pilicornis Gyllenhal				5		1				6
Unamis fulvipes Fall			1							1	3020
Tenebrionidae (2 species)
Helops pernitens LeConte									2	2
Scaphidema pictum LeConte				13						13	15
Throscidae (2 species)
Aulonthroscus validus LeConte				1						1
Pactopus hornii LeConte				2						2	3
Trogositidae (1 species)
Temnochila chlorodia Mannerheim				1						1	1
Zopheridae (1 species)
Phellopsis porcata LeConte					3					3	3